It may be asked how it could have occurred to any one, when pigeons,
for instance, first began to be domesticated, to wish to produce
fantails or pouters, since he could have no mental picture of them
in advance. Darwin replies to this objection, that it was not always
conscious and methodical artificial selection, such as is now
practised, that brought about the origin of breeds, but that they have
very often resulted, and at first perhaps always, from unconscious
selection. When savages tamed a dog, they used the 'best' of their dogs
for breeding, that is, they chose those which had in the highest degree
the qualities they valued, watchfulness, for instance, or if the dog
were intended for the chase, keen scent and swiftness. In this way the
body of the animal would be changed in a definite direction, especially
if rivalry helped, and if it was the ambition of each to possess a
dog as good as, or better than those of his tribal companions. That
perfectly definite changes in bodily form can thus be brought about
unconsciously is well illustrated by the case of a racehorse. This has
arisen within the last two hundred years simply because the fleetest
of the products of crossing between the Arab and the English horse
were always chosen for breeding. It could not have been predicted that
horses with thin neck, small head, long rump, and slender legs would
necessarily be the swiftest runners; but this is the form which has
resulted from the selection,--a very ugly, but very swift horse. This
unconscious selection must undoubtedly have played a large part in the
early stages of the evolution of the breeds of our domestic animals.
But even in the fully conscious and methodical selective breeding of
particular characters, the breeder rarely alters only the one his
attention is fixed on; generally quite a number of other characters
alter apart from his intention as an inevitable accompaniment of the
desired variation on which attention was riveted. There are breeds
of rabbits whose ears hang limply down instead of standing erect,
and in these so-called lop-eared rabbits the ear-muscles are partly
degenerated, and as a consequence of this lack of muscular strain the
skull has assumed another form. Thus the variation of one part may
influence the development of a second and a third organ, and may even
not stop there, for very often the influence has penetrated much deeper
and affected quite remote parts of the body.
Public-domain text, read in full here on John Shaqi.
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