Thus, although the fertility of each species is, as a matter of fact,
precisely regulated, a low rate of multiplication is not in itself
sufficient to prevent the excessive increase of any species, nor is
the quantity of the relevant food-supply. Whether this be very large
or very small, we see that in reality it is never entirely used up,
that, as a matter of fact, a much greater quantity is always left over
than has been consumed. If increase depended only on food-supply, there
would, for instance, be food enough in their tropical home for many
thousand times more elephants than actually occur; and among ourselves
the cockchafers might appear in much greater numbers than they do even
in the worst cockchafer year, for all the leaves of all the trees are
never eaten up; a great many leaves and a great many trees are left
untouched even in the years when the voracious insects are the most
numerous. Nor do the rose-aphides, notwithstanding their enormously
rapid multiplication, ever destroy all the young shoots of a rose-bush,
or all the rose-bushes of a garden, or of the whole area in which roses
grow.
At the same time it must be noted, that the number of individuals in
a species undoubtedly does bear some relation to the amount of the
food-supply available; for instance, it is very low among the large
carnivores, the lion, the eagle, and the like. In our Alps the eagles
have become rarer with the decrease of game, and where one eagle pair
make their eyrie they rule alone over a hunting territory of more than
sixty miles, a preserve on which no others of the same species are
allowed to intrude. If there were several pairs of eagles in such a
preserve, they would soon have so decimated the food-supply that they
would starve. On the other hand, numerous herbivores, e.g. chamois and
marmots, live within the bounds of the pair of eagles' hunting grounds,
since the food they require is present in enormously greater quantity.
Public-domain text, read in full here on John Shaqi.
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