When the number of foxes in a hunting district increases, the number of
the hares that they prey upon diminishes, and, on the other hand, the
decimating of the foxes by Man brings about an increase in the number
of hares in the district. Under natural conditions, that is, without
the intervention of Man, there would be a constant balancing of the
numbers of hares and foxes, for every noteworthy increase of the hares
would be followed by a similar increase of foxes, and this, in its
turn, would diminish the number of hares, so that they would no longer
suffice for the support of so many foxes, and these would decrease in
number again, until the number of hares had again increased because
of the lessened persecution and elimination. In nature the case is not
quite so simple, because the fox does not live on hares alone, and the
hare is not preyed upon only by the fox; but the illustration may serve
to elucidate the point that a moving equilibrium is maintained between
the species of a district, between persecutors and persecuted, in such
a way that the number of individuals in the two species is always
varying a little up and down, and that each influences the other so
that a regulative process results. Throughout periods of considerable
length the average remains the same; that is to say, a _normal number_
is established. This normal strength of population is the mean above
and below which the number of individuals is constantly varying. It is,
of course, seldom that the mutual influences and regulations are so
simple as in the example given; usually several or even many species
interact upon each other, and not beasts of prey and their victims
alone, but the most diverse species of animals and plants, which do
not stand in any obvious relation to one another at all. Moreover, the
physical, and especially the climatic conditions, also cause the normal
number of the species to rise and fall.
The inter-relations between species living together on the same area
are so intricate that I should like to give two other illustrations.
Let us first take Darwin's famous instance of the fertility of clover,
which depends on the number of cats. It is of course only an imaginary
one, but the facts it is based upon are quite correct. The number of
cats living in a village to a certain extent determines the number of
field-mice in the neighbourhood. These again destroy the nests of the
humble-bees, which live in holes in the ground, and thus the number
of humble-bees depends on that of the field-mice and cats. But the
clover must be pollinated by insects if it is to produce fertile seed,
and only the humble-bee has a proboscis long enough to effect the
pollination. Therefore the quantity of clover-seed annually produced
depends on the number of humble-bees, and ultimately upon the number of
cats. And, as a matter of fact, humble-bees were introduced into New
Zealand from England, because without them the clover would produce no
fertile seeds.
Public-domain text, read in full here on John Shaqi.
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