A modern opponent of the selection theory (Eimer) has suggested
that the marking of the lateral veins, and other resemblances to a
leaf in _Kallima_, represent nothing more than the pattern which
was present in any case, inherited from ancestors, and which in the
course of time arranged itself in a particular manner according to
internal developmental laws. Not selection--that is, adaptation to
surroundings--but the internal developmental impulse has brought about
the resemblance to the leaf. It is astonishing how a preconceived
idea can blind a man and weaken his judgment! It goes without saying
that the adaptations do not start from a _tabula rasa_, but from
what is already present; of course, natural selection makes use of
the markings inherited from ancestors; it takes what already exists,
and alters or extends it as suits best. Thus it is easy to prove
that the clear mirrors (Fig. 13, _gl_^1 and _gl_^2) on the wings of
_Kallima_ have arisen from a modification of the nuclei of eye-spots,
just as the dark mould-spots which often occur, frequently develop
in association with the inherited eye-spots; not always however, for
many such accumulations of black scales occur in spots on which there
has never been an eye-spot. Thus, too, the 'midribs' of the butterfly
have in part arisen from a gradual displacing, extending, and altering
of the direction of inherited stripes as, for instance, is clearly
recognizable in the posterior wing of Fig. 13, but sometimes they are
new formations. But the veining of a leaf is never found on the wing
of any butterfly of a species which has not the habit of resting among
leaves, or which has not had it at one time, and it never corresponds
to the natural marking of any genus which does not live in forests.
This impression of leaf-venation has obviously arisen from quite
different patterns of markings, and it has been reached now by one
way, now by another. We can see this from the fact that, in different
butterflies, it lies in quite different positions on the wing. In
the _Kallima_ species the stalk of the leaf lies in the tail of the
posterior wing, the tip of the midrib lies near the tip of the wing; in
_Cœnophlebia archidona_ it is exactly reversed, the tip of the anterior
wing (Fig. 14) is prolonged, and forms the stalk, while a broad,
dark, stripe, the midrib (_mr_), runs from there across the middle of
both wings, and seems to give off two or three lateral ribs running
outwards. If it be asked whether this butterfly always sits down so
artistically that the 'upward turning leaf-stalk is in juxtaposition
to a twig,' we may answer that a bird flying fast is not likely to
look to see whether every leaf in the profusion of foliage in the
primitive forests is properly fastened to its stalk or not, any more
than we should do in the case of a painted bush, on which many a leaf
has the appearance of floating in the air, just as in nature, or in its
faithful copy, the photograph.
Public-domain text, read in full here on John Shaqi.
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