This is certainly ingeniously and cleverly thought out, and it reads
even better and more smoothly in the original than in my brief summary,
but it will hardly be regarded as a refutation of my position; the
hypotheses are all too daring for that. We have no knowledge that
particular modifications in form can be produced and conditioned by
particular kinds of food, and, indeed, the contrary has been proved,
namely, that the two or three different castes of polymorphic species
have precisely similar diet. I need only recall the six forms of female
in _Papilio merope_, of which at least three have been obtained from
the same set of eggs, and by feeding with the same plant.
It is true that there are ants which lay in stores of nourishment,
but these consist, for the most part, of one kind of seeds, or of
honey, not of different substances, and we have no knowledge that
the different persons use different food, or even that there is any
diversity in the mode of feeding the helpless larvæ. The feeding in
some species takes place from mouth to mouth, and therefore cannot be
precisely investigated, and we can only suppose from analogy with bees
that the larvæ of the males and females frequently receive not only
more abundant, but qualitatively different food. They are fed from the
crop unless the food consists of the pith of a tree in which the larvæ
are imbedded, as Dahl informs us is the case with some tropical ants of
the Bismarck Archipelago.
But even if we assume that the soldiers take different food from the
ordinary workers, and different again from that of the sexual animals,
is it by virtue of the quality of their food that they have become
what they are? Have our breeds of pigeons or hens been produced by
different diet, or do we know anything in the whole range of animal
life of such a parallelism between food and bodily structure as Zehnder
here assumes? And if, in reality, let us say, the breeds of pigeon had
arisen through specific dieting, and we were to feed one pair with the
specific food-stuffs of three different breeds, would the descendants
of this pair exhibit the form of these three breeds? Or would they
exhibit them in precisely the proportion in which the food-stuffs had
been mixed? It seems to me that Zehnder's assumptions diverge so far
from what we are accustomed to regard as solid ground in biology that
they hardly require consideration, and yet he not only uses them for
the explanation of the case of the ants, but bases upon them the whole
of his theory of the inheritance of acquired characters.
Public-domain text, read in full here on John Shaqi.
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