It does not affect matters to discover cases in which we cannot
recognize this relation between the regenerative capacity of a part
and its importance or its liability to injury. Such instances do not
lessen the convincingness of the positive cases, since we do not know
the exact conditions which may lead to the increase of regenerative
capacity in a part, and, above all, since we do not know the rate at
which such an increase may take place. If adaptation in general depends
upon processes of selection, these processes must also be able to give
rise to an increase in the power of regeneration. On the other hand, it
by no means follows that the disappearance of a faculty of regeneration
which was once present in a part, but which has become superfluous
in the course of time, must take place immediately through natural
selection. For it is the very essence of natural selection that it only
furthers what is useful, and only removes what is injurious; over what
is indifferent it has no power at all. Thus it follows that the faculty
of regeneration, when it has once been present in a part, cannot be
set aside by natural selection (personal selection), for it is in no
way injurious to its possessor. If it gradually decreases and becomes
extinct notwithstanding this, when it is of no further use, as seems to
be to some extent the case in regard to the legs and tail of the blind
Proteus, that must depend on other processes, on those which generally
bring about the gradual disappearance of disused parts or capacities.
We shall attempt to probe to the roots of these processes later on;
for the present let it suffice us to know that, according to our
experience, they go on with exceeding slowness, and that it has taken
whole geological periods to eliminate the legs of the snake-ancestors
so completely as has been done from the structure of most of our modern
snakes, while the Proteus which migrated into the caves of Krain as far
back as the Cretaceous period is indeed blind, but still retains its
eyes under the skin, though in a degenerate condition.
Public-domain text, read in full here on John Shaqi.
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