In most species these ring-spots occur on many segments (10-12) (Fig.
117, _B_), and in cases where they are of importance in making the
caterpillar conspicuous and easily seen they sometimes form a double
row. But we know one species, _Deilephila hippophaës_, in which only
a single ring-spot exists, and it is a large brick-red spot on the
second last segment, mimicking the red berry of the buckthorn (Fig.
8, _A_ and _B_, _r_). But individuals also occur in which there are,
on the five or six segments in front, smaller ring-spots which become
less distinct the further forward they are, and in most caterpillars
it is possible, on careful examination, to recognize little red dots
on the faded sub-dorsal stripes of these segments (Fig. 8, _B_). We
might be disposed to think, on this account, that the ancestors of
_D. hippophaës_ bore rings on all the segments, and that these had
gradually become vestigial on the majority of them, because they had
lost their earlier biological importance, and now, by adaptation to the
buckthorn, could only be of use on the second last. But when we take
the ontogeny also into account we find in the young caterpillar only a
simple sub-dorsal line, upon which, in the third stage, the red spot of
the tail-horn segment appears (Fig. 8, _A_).
No spots ever occur on the other segments at this stage; they only
appear in the last stage, but as they may be entirely wanting, they
must have arisen as the result of internal laws of correlation, that
is, they must be recapitulations of the hindmost spots which arose
in the phylogeny through natural selection. We may conclude this, at
least, if we believe in the truth of the fundamental proposition of the
biogenetic law, and admit that there is in the ontogeny some more or
less distinct recapitulation of the phylogeny.
[Illustration: FIG. 118. Two stages in the life-history of the Spurge
Hawk-moth (_Deilephila euphorbiæ_). _A_, first stage, the caterpillar
dark blackish-green, without marking. _B_, second stage, the row of
spots is distinctly connected by a light streak, the vestige of the
sub-dorsal stripe.]
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