This proposition may be recognized as true in the case of _Deilephila_
also, if we compare the different species with one another as regards
their ontogeny. We find here too that not only the sub-dorsal, that is,
the phyletically oldest marking of the Sphingid caterpillars, occurs
everywhere in the young stages, but also that it is being shunted back
to younger and younger stages, in proportion to the degree of the
development of the spot-marking reached in the full-grown caterpillar.
Thus, for instance, in the caterpillar of _Deilephila euphorbiæ_ the
highest form of spot-marking is reached, and in this species the
sub-dorsal line is no longer the sole marking element at any stage.
Leaving out of the question the absolutely unmarked little caterpillar
which emerges from the egg (Fig. 118, _A_), there appears at once in
the second stage a series of ring-spots connected by a fine white
sub-dorsal line (Fig. 118, _B_). In the following stage, the third,
this sub-dorsal line disappears without leaving a trace, and there
remains only the spot-marking, which is subsequently duplicated.
Let us compare with this the ontogeny of the bed-straw hawk-moth,
_Deilephila galii_ (Fig. 117). The full-grown caterpillar possesses
only a single row of ring-spots (_B_), and accordingly the young stages
of the caterpillar up to the fourth show a distinct sub-dorsal line
(_A_), although spots are seen upon it. A still earlier phyletic stage
of development is illustrated by _Deilephila livornica_, in which the
ring-spots are all connected by the sub-dorsal line.
It can thus hardly be doubted that the biogenetic law is guiding us
aright when we conclude from a comparison of the ontogeny of the
different species of _Deilephila_, that the oldest ancestors of the
genus possessed only the longitudinal stripes, and that from these
small pieces were cut off as ring-spots, and that these were gradually
perfected and ultimately duplicated, while at the same time the
original marking, the longitudinal stripe, was shunted back further and
further in the young stages, until it finally disappeared altogether.
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