Therefore, when a species is most perfectly adapted to its conditions,
it would of necessity begin to degenerate if personal selection were
not continually guarding it, and setting aside everything that is in
excess or deficient as soon as it begins to be prejudicial. But the
adaptation of a species does not depend upon _one_ character persisting
at its normal level, but on the persistence of very many, and many of
these vary simultaneously upwards or downwards, and reach the limit of
selection-value at one time or another. If there were no amphimixis,
then either all individuals with any excessive variant would be at
once eliminated, or the species would go on deteriorating until this
excessive variant was so numerously and strongly represented in all
its individuals that it would perish through degeneration. But even
in the first of these cases the species would drift towards the fate
of extinction, because excessive variations do occur even in every
asexual generation, and would appear in an increasingly large number
of determinants if there were no possibility of rejecting them and
eliminating them from the lineage of the species.
This is made possible through the periodic intervention of amphimixis;
it is actually effected thereby; and in this way alone the species
is kept at its high-water mark of adaptation. It is not necessary
to assume that every single determinant which is varying in an
unfavourable direction is at once eliminated as soon as it becomes
prejudicial, that is, reaches negative selection-value, or--to make
use of an expression introduced by Ammon--as soon as it oversteps the
boundaries of the 'playground of variations,' the limits within which
variations are neither favourable nor unfavourable. But _in the course
of generations_ they are unfailingly eliminated, especially when a
large number of unfavourably varying determinants are coincident in
the germ-plasm. Then the individuals which arise from a germ-plasm
thus composed must perish in the struggle for existence, and thus the
id-combinations with excessive determinants are eliminated from the
germinal constitution of the species. As this is repeated as often as
excesses of the ids occur, the species is kept pure.
It might be objected that, through such a continual weeding-out of
rebellious determinants, the germ-plasm would become so constant in
its constitution that it would ultimately be secure from all such
aberrations of it on the part of its determinants, and therefore would
in time become quite incapable of diverging from its proper path at
all, and would thus no longer require this continual correction through
amphimixis.
Public-domain text, read in full here on John Shaqi.
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