Let us return after this digression to our previous train of thought;
we have to inquire how we can interpret the fact of continued
self-fertilization without any visible injurious results to the
species. If cross-fertilization be a material advantage as regards the
continuance of the species, how can it be transformed into its opposite
without evil effects? And there are no visible evil effects in _Ophrys
apifera_. It is indeed not so abundant as _Ophrys muscifera_, or other
allied species, but it certainly does not follow from that that it is
on the way to extinction; certainly no decrease either of vigour of
growth or of fertility can be observed.
If we inquire from the standpoint of our theory, how the composition
of the germ-plasm must have altered through continual inbreeding,
we have already found the answer--that through the reduction of the
number of ids at the maturation of every germ-cell the diversity of the
germ-plasm would gradually be lessened, that the number of different
ids would thereby be lessened possibly even to the identity of the
whole of the ids.
The consequences of such extreme uniformity of the germ-plasm would
not, according to our theory, necessarily be that the species would
be incapable of continued existence, but it would be that the species
would become incapable of adaptations in many directions. Adaptations
in one direction, such, for instance, as the variation in the mode of
attachment and detachment of the pollinia of an Orchid, would still be
possible. Thus a species which has long been perfectly adapted will be
able to make the transition to inbreeding without injury to its chances
of continued existence, if it be compelled by circumstances to do so.
Species, on the other hand, which are still undergoing considerable
transformations in many directions must be exposed by these to the
danger of degeneration, just as happens in the artificial experiments
with domesticated animals, whose secret weaknesses are greatly
exaggerated by inbreeding.
We might be inclined to regard the effects of inbreeding as similar to
those of parthenogenesis; they are certainly analogous, for both modes
of reproduction must lead to a certain degree of uniformity in the
germ-plasm. But there seems to me to be a difference and one which is
not without importance.
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