it is rather a purely mechanical process, which takes its course quite
independently of what is useful or disadvantageous, and that it must
take this course according to the given regeneration-mechanism and the
stimulus supplied in the special case. It cannot be supposed that these
supernumerary heads and tails are purposeful, but who would expect an
adaptive reaction from the animal in a case like this, since cuts of
the kind which we make artificially, and _must keep open artificially_
if the deformities are to develop, hardly occur in nature, and, if they
did occur, would very quickly close up again? Adaptations can only
develop in response to conditions which occur and recur in a majority
of cases, and when they have a useful, that is, species-preserving
result. The adaptiveness of the organism is blind, it does not see the
individual case, it only takes into account the cases in the mass,
and acts as it must after the mechanism has once been evolved. The
case is the same as that of 'aberrant' or mistaken instincts, whose
origin by means of selection is the more clearly proved, since we must
recognize such an instinct as a pure mechanism and not as the outcome
of purposeful forces.
In the regeneration of Planarians we must think of the
regeneration-idioplasm as containing the full complex of the
collective determinants of the three germinal layers, and possibly
we must add to this cells with the complete germ-plasm for giving
rise to the reproductive cells. But when the amputated tail of the
newt is regenerated, or its leg, or the arm of a starfish, or the
bill of a bird, we have no ground for assuming that the cells, from
which regeneration starts, contain the whole germ-plasm, since the
determinants of the replaceable parts suffice to explain the facts.
We must even dispute the possibility of the presence of the whole
germ-plasm in this case, because the faculty of regeneration of the
relevant cells is really no longer a general one, but is limited to
the reproduction of a particular part. This is seen in the fact that,
in the starfish, whose high regenerative capacity is well known, the
central disk of the body may indeed give rise to new arms[2]; but
an excised arm, to which no part of the disk adheres, is in most
starfishes unable to give rise to the body. Thus the arm does not
contain in its cells the determinants of the disk, but the latter
contains those of the arm. We are not surprised that the amputated
tail of the salamander does not reproduce the whole animal, but this
can only be because the impelling forces to the regeneration of the
whole animal are wanting, that is, that the cut surface only contains
the determinants of the tail and not the complete germ-plasm. It might
be objected here that the tail-piece is too small to give rise to the
whole body, but in _Planaria_ it is only very diminutive heads and
tails which grow from the artificial incisions, and the same is true
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