that _special_ 'primary constituents' are necessary, without which the
relevant part cannot arise. The indispensableness of these primary
constituents, the determinants, seems to me to depend on the fact
that the new structure cannot be built up simply by procuring organic
material, but _that specially hewn stones, different in every case,
are necessary_, which can only be supplied in virtue of an historical
transmission, or, to abandon the metaphor, because the vital units of
which the organ is to be reconstructed possess a specific character
and have a long history behind them; thus they can only arise from
such vital units as have been handed on through generations, that is,
from the determinants. But these primary constituents are given to the
different forms of life in very varying degrees and in very unequal
distribution, and as far as we can see according to their suitability
to an end.
LECTURE XXII
SHARE OF THE PARENTS IN THE BUILDING UP OF THE OFFSPRING
The ids are 'ancestral plasms'--The reducing division brings
about a diversity of germ-plasm in the germ-cells--Bolles
Lee's 'Neotaxis' even in the primordial germ-cells--Häcker's
observations on the persistent distinctness of the maternal and
paternal chromosomes--Identical twins--The individuality is
determined at fertilization--Unequal share of the ids in the
determination of the offspring--Preponderance of one parent in the
composition of the offspring--Certain ids of the ancestors remain
unchanged in the germ-plasm of the descendants--Struggle of the
Biophors--Alternation of the hereditary sequences in the parts
of the child--Reversion--Datura-hybrids--Zebra-striping in the
horse--Three-toed horses--New experiments in hybridization among
plants by Correns and De Vries--Xenia.
As far as the phenomena of regeneration and budding are concerned, we
have not been able to do much more than bring them under a formula,
which harmonizes with the germ-plasm theory. But the case is different
with the actual phenomena of inheritance in the restricted sense, for
instance, with regard to the transmission of individual peculiarities
_from parent to child_. Here the theory really increases our insight
and lets us penetrate deeper into the causes of the phenomena; it is
here no longer a mere 'portmanteau-theory.'
Public-domain text, read in full here on John Shaqi.
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