If the germ-plasm consists of ids, these ids collectively must
determine the structure, the whole individuality--let us say, briefly,
the 'type' of the offspring; it is the resultant of all the different
impelling forces which are contained in the different ids. If these
were all equally strong, and all operating in the same direction,
they would necessarily all have the same share in the resultant of
development, the 'type' of the child. But this is not the case.
Numerous experiments on the hybridization of two species of plant have
taught us that the descendants of such hybridization usually maintain
a medium between the ancestral species; but it is not always the case,
for in many hybrids the character of one species, whether paternal or
maternal, preponderates in the young plant.
We recognize the same thing still more clearly in Man, whose children
by no means always maintain a medium between the characters of the two
parents, but frequently resemble one--the father or the mother--much
more strongly than the other.
How can this fact be theoretically explained? Must we ascribe to the
ids of the father or of the mother a greater determining power? Without
excluding such an assumption as on _a priori_ grounds inadmissible,
I am inclined to believe that we do not require it to explain _this_
phenomenon. For, if we take our stand simply on the fact of the
preponderance of one parent, it follows directly from this that not
all the ids control the type of the child, let the cause of the
non-co-operation of some of them be what it may. But if in this case
only a portion of the ids contained in the germ-plasm controls the
type, this combination of ids suffices to make the child resemble
one parent, the father, for instance, and consequently half the
number of ids is sufficient in some circumstances to determine the
child--taking for granted that the one-sidedness of the inheritance is
complete, which never actually happens. But the half number of ids can
only suffice if it includes the same combinations of ids which have
determined the type in the case of the father; as soon as one or more
ids of this particular combination are replaced by others the paternal
germ-plasm alone is not enough to call forth complete resemblance in
the child.
But, at the reduction, a change of arrangement of ids takes place,
and a new combination arises, and thus each germ-cell receives its
particular group of ids. It may thus happen that, in one particular
sperm-cell, exactly the same group of ids is contained as that which
determined the type of the father, and that the same is true of a
particular egg-cell in regard to the type of the mother. Let us now
assume that a sperm-cell and an egg-cell meet, which contain both those
groups of ids which had determined the type of the father and of the
mother; if the determining power of the maternal and paternal ids were
equal a child would result which would maintain the medium between
father and mother.
Public-domain text, read in full here on John Shaqi.
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