[12] Morton, 'The Problem of Heredity in Reference to Inebriety,'
Proceed. Soc. for the Study of Inebriety, No. 42, Nov. 1894.
But even if there were not a trace of proof of the transmissibility of
functional modifications, that alone would not justify us in concluding
that the transmission is impossible, for many things may happen that we
are not in a position to prove at present. If it could be shown that
there was a whole group of phenomena that could not be explained in any
other way than on the hypothesis of such inheritance, then we should
be obliged to assume that it really occurred, although it was not
demonstrable, and, indeed, not even theoretically conceivable. This is
the standpoint of the adherents of the Lamarckian principle at present.
They say there are a great number of transformations which are simply
and easily explained, if we regard them as the effects of inherited
use or disuse, but which admit only of a strained explanation, and
sometimes of none at all, on the basis of natural selection, and these
are not a few isolated cases, but whole categories of them.
I will submit a few of these, and show at the same time why I cannot
regard them as convincing, even if it be the case that we are not at
present in a position to explain them without the aid of the Lamarckian
principle. But let me hasten to add that it is my belief that we can do
this, although certainly not without first giving a somewhat extended
application to the principle of selection.
It has often been maintained that the existence of animal instincts is
in itself enough to prove that the Lamarckian principle is operative.
In one of the earlier lectures I showed that at least the greater
number of instincts must have originated in purely reflex actions,
and therefore, like these actions themselves, can only be explained
through natural selection. A reflex action, such as coughing, sneezing,
shutting of the eyelids, and so on, differs from an instinctive
action in the lesser complexity and shorter duration of the series
of movements liberated by a sense-impression, and also in that it
does not require to enter into consciousness at all; but no very
precise boundary can be drawn between the two, and, in any case, both
depend, as we have already seen, on a quite analogous anatomical
basis. It is only a difference in degree whether, at the sight of a
rapidly approaching object, the muscles of the eyelids contract, and
by shutting the lids, protect the eye, or whether the fly, which we
intend to seize with our hand, is impelled by the sight of the rapidly
approaching shadow of the hand to fly quickly up. The action of the fly
may be regarded as reflex, or equally well as instinctive. But there
is also only a difference in degree, not in kind, between this simple
action and the complex and protracted behaviour of a mason-bee, the
sight of whose colony impels her to fly out and fetch clay, with it
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