upon stray specimens if you search carefully in damp places under
fallen leaves. Even in testacella, however, the small shell is still
external. In this yellow slug here, on the contrary, it does not show
itself at all, but is buried under the closely wrinkled skin of the
glossy mantle. It has become a mere saucer, with no more symmetry or
regularity than an oyster-shell. Among the various kinds of slugs, you
may watch this relic or rudiment gradually dwindling further and
further towards annihilation; till finally, in the great fat black
slugs which appear so plentifully on the roads after summer showers, it
is represented only by a few rough calcareous grains, scattered up and
down through the mantle; and sometimes even these are wanting. The
organs which used to secrete the shell in their remote ancestors have
either ceased to work altogether or are reduced to performing a useless
office by mere organic routine.
The reason why some mollusks have thus lost their shells is clear
enough. Shells are of two kinds, calcareous and horny. Both of them
require more or less lime or other mineral matters, though in varying
proportions. Now, the snails which thrive best on the bare chalk downs
behind my little combe belong to that pretty banded black-and-white
sort which everybody must have noticed feeding in abundance on all
chalk soils. Indeed, Sussex farmers will tell you that South Down
mutton owes its excellence to these fat little mollusks, not to the
scanty herbage of their thin pasture-lands. The pretty banded shells in
question are almost wholly composed of lime, which the snails can, of
course, obtain in any required quantity from the chalk. In most
limestone districts you will similarly find that snails with calcareous
shells predominate. But if you go into a granite or sandstone tract you
will see that horny shells have it all their own way. Now, some snails
with such houses took to living in very damp and marshy places, which
they were naturally apt to do--as indeed the land-snails in a body are
merely pond-snails which have taken to crawling up the leaves of
marsh-plants, and have thus gradually acclimatised themselves to a
terrestrial existence. We can trace a perfectly regular series from the
most aquatic to the most land-loving species, just as I have tried to
trace a regular series from the shell-bearing snails to the shell-less
slugs. Well, when the earliest common ancestor of both these last-named
races first took to living above water, he possessed a horny shell
(like that of the amber-snail), which his progenitors used to
manufacture from the mineral matters dissolved in their native streams.
Some of the younger branches descended from this primæval land-snail
took to living on very dry land, and when they reached chalky districts
manufactured their shells, on an easy and improved principle, almost
entirely out of lime. But others took to living in moist and boggy
Public-domain text, read in full here on John Shaqi.
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