The arum, like the grain of wheat, has only a single seed-leaf; whereas
the pea, as we all know, has two. This is the most fundamental
difference among flowering plants, as it points back to an early and
deep-seated mode of growth, about which they must have split off from
one another millions of years ago. All the one-lobed plants grow with
stems like grasses or bamboos, formed by single leaves enclosing
another; all the double-lobed plants grow with stems like an oak,
formed of concentric layers from within outward. As soon as the arum,
with its sprouting head, has raised its first leaves far enough above
the ground to reach the sunlight, it begins to form fresh starches and
new leaves for itself, and ceases to be dependent upon the store laid
up in its buried lobe. Most seeds accordingly contain just enough
material to support the young seedling till it is in a position to
shift for itself; and this, of course, varies greatly with the habits
and manners of the particular species. Some plants, too, such as the
potato, find their seeds insufficient to keep up the race by
themselves, and so lay by abundant starches in underground branches or
tubers, for the use of new shoots; and these rich starch receptacles we
ourselves generally utilise as food-stuffs, to the manifest detriment
of the young potato-plants, for whose benefit they were originally
intended. Well, the arum has no such valuable reserve as that; it is
early cast upon its own resources, and so it shifts for itself with
resolution. Its big, glossy leaves grow apace, and soon fill out, not
only with green chlorophyll, but also with a sharp and pungent essence
which makes them burn the mouth like cayenne pepper. This acrid juice
has been acquired by the plant as a defence against its enemies. Some
early ancestor of the arums must have been liable to constant attacks
from rabbits, goats, or other herbivorous animals, and it has adopted
this means of repelling their advances. In other words, those arums
which were most palatable to the rabbits got eaten up and destroyed,
while those which were nastiest survived, and handed down their
pungency to future generations. Just in the same way nettles have
acquired their sting and thistles their prickles, which efficiently
protect them against all herbivores except the patient, hungry donkey,
who gratefully accepts them as a sort of _sauce piquante_ to the
succulent stems.
Public-domain text, read in full here on John Shaqi.
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