Behind the old mill, whose overshot wheel, backed by a wall thickly
covered with the young creeping fronds of hart's-tongue ferns, forms
such a picturesque foreground for the view of our little valley, the
mill-stream expands into a small shallow pond, overhung at its edges by
thick-set hazel-bushes and clambering honeysuckle. Of course it is only
dammed back by a mud wall, with sluices for the miller's water-power;
but it has a certain rustic simplicity of its own, which makes it
beautiful to our eyes for all that, in spite of its utilitarian origin.
At the bottom of this shallow pond you may now see a miracle daily
taking place, which but for its commonness we should regard as an
almost incredible marvel. You may there behold evolution actually
illustrating the transformation of life under your very eyes: you may
watch a low type of gill-breathing gristly-boned fish developing into
the highest form of lung-breathing terrestrial amphibian. Nay,
more--you may almost discover the earliest known ancestor of the whole
vertebrate kind, the first cousin of that once famous ascidian larva,
passing through all the upward stages of existence which finally lead
it to assume the shape of a relatively perfect four-legged animal. For
the pond is swarming with fat black tadpoles, which are just at this
moment losing their tails and developing their legs, on the way to
becoming fully formed frogs.
The tadpole and the ascidian larva divide between them the honour of
preserving for us in all its native simplicity the primitive aspect of
the vertebrate type. Beasts, birds, reptiles, and fishes have all
descended from an animal whose shape closely resembled that of these
wriggling little black creatures which dart up and down like imps
through the clear water, and raise a cloud of mud above their heads
each time that they bury themselves comfortably in the soft mud of the
bottom. But while the birds and beasts, on the one hand, have gone on
bettering themselves out of all knowledge, and while the ascidian, on
the other hand, in his adult form has dropped back into an obscure and
sedentary life--sans eyes, sans teeth, sans taste, sans everything--the
tadpole alone, at least during its early days, remains true to the
ancestral traditions of the vertebrate family. When first it emerges
from its egg it represents the very most rudimentary animal with a
backbone known to our scientific teachers. It has a big hammer-looking
head, and a set of branching outside gills, and a short distinct body,
and a long semi-transparent tail. Its backbone is a mere gristly
channel, in which lies its spinal cord. As it grows, it resembles in
every particular the ascidian larva, with which, indeed, Kowalewsky and
Professor Ray Lankester have demonstrated its essential identity. But
since a great many people seem wrongly to imagine that Professor
Lankester's opinion on this matter is in some way at variance with Mr.
Darwin's and Dr. Haeckel's, it may be well to consider what the
Public-domain text, read in full here on John Shaqi.
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