species in animals so very variable as fish. There are, in fact, no
less than twelve kinds of trout wholly peculiar to the British Islands,
and some of these are found in very restricted areas. Thus, the Loch
Stennis trout inhabits only the tarns of Orkney; the Galway sea trout
lives nowhere but along the west coast of Ireland; the gillaroo never
strays out of the Irish loughs; the Killin charr is confined to a
single sheet of water in Mayo; and other species belong exclusively to
the Llanberis lakes, to Lough Melvin, or to a few mountain pools of
Wales and Scotland. So great is the variety that may be produced by
small changes of food and habitat. Even the salmon himself is only a
river trout who has acquired the habit of going down to the sea, where
he gets immensely increased quantities of food (for all the trout kind
are almost omnivorous), and grows big in proportion. But he still
retains many marks of his early existence as a river fish. In the first
place, every salmon is hatched from the egg in fresh water, and grows
up a mere trout. The young parr, as the salmon is called in this stage
of its growth, is actually (as far as physiology goes) a mature fish,
and is capable of producing milt, or male spawn, which long caused it
to be looked upon as a separate species. It really represents, however,
the early form of the salmon, before he took to his annual excursion to
the sea. The ancestral fish, only a hundredth fraction in weight of his
huge descendant, must have somehow acquired the habit of going
seaward--possibly from a drying up of his native stream in seasons of
drought. In the sea, he found himself suddenly supplied with an
unwonted store of food, and grew, like all his kind under similar
circumstances, to an extraordinary size. Thus he attains, as it were,
to a second and final maturity. But salmon cannot lay their eggs in the
sea; or at least, if they did, the young parr would starve for want of
their proper food, or else be choked by the salt water, to which the
old fish have acclimatised themselves. Accordingly, with the return of
the spawning season there comes back an instinctive desire to seek once
more the native fresh water. So the salmon return up stream to spawn,
and the young are hatched in the kind of surroundings which best suit
their tender gills. This instinctive longing for the old home may
probably have arisen during an intermediate stage, when the developing
species still haunted only the brackish water near the river mouths;
and as those fish alone which returned to the head waters could
preserve their race, it would soon grow hardened into a habit engrained
in the nervous system, like the migration of birds or the clustering of
swarming bees around their queen. In like manner the Jamaican
land-crabs, which themselves live on the mountain-tops, come down every
year to lay their eggs in the Caribbean; because, like all other crabs,
they pass their first larval stage as swimming tadpoles, and afterwards
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