so as finally to evolve conduct which differs very widely with
different circumstances and different characters. Even though it be
true, as determinists believe (and I reckon myself among them), that
such conduct is the necessary result of a given character and given
circumstances--or, if you will, of a particular set of nervous
structures and a particular set of external stimuli--yet we all know
that it is capable of varying so indefinitely, owing to the complexity
of the structures, as to be practically incalculable. But it is not so
with the butterfly. His whole life is cut out for him beforehand; his
nervous connections are so simple, and correspond so directly with
external stimuli, that we can almost predict with certainty what line
of action he will pursue under any given circumstances. He is, as it
were, but a piece of half-conscious mechanism, answering immediately to
impulses from without, just as the thermometer answers to variations of
temperature, and as the telegraphic indicator answers to each making
and breaking of the electric current.
In early life the future butterfly emerges from the egg as a
caterpillar. At once his many legs begin to move, and the caterpillar
moves forward by their motion. But the mechanism which set them moving
was the nervous system, with its ganglia working the separate legs of
each segment. This movement is probably quite as automatic as the act
of sucking in the new-born infant. The caterpillar walks, it knows not
why, but simply because it has to walk. When it reaches a fit place for
feeding, which differs according to the nature of the particular larva,
it feeds automatically. Certain special external stimulants of sight,
smell, or touch set up the appropriate actions in the mandibles, just
as contact of the lips with an external body sets up sucking in the
infant. All these movements depend upon what we call instinct--that is
to say, organic habits registered in the nervous system of the race.
They have arisen by natural selection alone, because those insects
which duly performed them survived, and those which did not duly
perform them died out. After a considerable span of life spent in
feeding and walking about in search of more food, the caterpillar one
day found itself compelled by an inner monitor to alter its habits.
Why, it knew not; but, just as a tired child sinks to sleep, the gorged
and full-fed caterpillar sank peacefully into a dormant state. Then its
tissues melted one by one into a kind of organic pap, and its outer
skin hardened into a chrysalis. Within that solid case new limbs and
organs began to grow by hereditary impulses. At the same time the form
of the nervous system altered, to suit the higher and freer life for
which the insect was unconsciously preparing itself. Fewer and smaller
ganglia now appeared in the tail segments (since no legs would any
longer be needed there), while more important ones sprang up to govern
the motions of the four wings.
Public-domain text, read in full here on John Shaqi.
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