“Finally, as indefinite and almost illimitable variability is the
usual result of domestication and cultivation, with the same part or
organ varying in different individuals in different or even in
directly opposite ways; and as the same variation, if strongly
pronounced, usually recurs only after long intervals of time, any
particular variation would generally be lost by crossing, reversion,
and the accidental destruction of the varying individuals, unless
carefully preserved by man.”—Vol. ii, 292.
Remembering that mankind, subject as they are to this domestication and
cultivation, are not, like domesticated animals, under an agency which
picks out and preserves particular variations; it results that there
must usually be among them, under the influence of natural selection
alone, a continual disappearance of any useful variations of particular
faculties which may arise. Only in cases of variations which are
specially preservative, as for example, great cunning during a
relatively barbarous state, can we expect increase from natural
selection alone. We cannot suppose that minor traits, exemplified among
others by the æsthetic perceptions, can have been evolved by natural
selection. But if there is inheritance of functionally-produced
modifications of structure, evolution of such minor traits is no longer
inexplicable.
* * * * *
Two remarks made by Mr. Darwin have implications from which the same
general conclusion must, I think, be drawn. Speaking of the variability
of animals and plants under domestication, he says:—
“Changes of any kind in the conditions of life, even extremely
slight changes, often suffice to cause variability.... Animals and
plants continue to be variable for an immense period after their
first domestication; ... In the course of time they can be
habituated to certain changes, so as to become less variable; ...
There is good evidence that the power of changed conditions
accumulates; so that two, three, or more generations must be exposed
to new conditions before any effect is visible.... Some variations
are induced by the direct action of the surrounding conditions on
the whole organization, or on certain parts alone, and other
variations are induced indirectly through the reproductive system
being affected in the same manner as is so common with organic
beings when removed from their natural conditions.”—(_Animals and
Plants under Domestication_, vol. ii, 270.)
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