Out of this dead lock we were released—in large measure, though not I
believe entirely—by the _Origin of Species_. That work brought into view
a further factor; or rather, such factor, recognized as in operation by
here and there an observer (as pointed out by Mr. Darwin in his
introduction to the second edition), was by him for the first time seen
to have played so immense a part in the genesis of plants and animals.
Though laying myself open to the charge of telling a thrice-told tale, I
feel obliged here to indicate briefly the several great classes of facts
which Mr. Darwin's hypothesis explains; because otherwise that which
follows would scarcely be understood. And I feel the less hesitation in
doing this because the hypothesis which it replaced, not very widely
known at any time, has of late so completely dropped into the
background, that the majority of readers are scarcely aware of its
existence, and do not therefore understand the relation between Mr.
Darwin's successful interpretation and the preceding unsuccessful
attempt at interpretation. Of these classes of facts, four chief ones
may be here distinguished.
In the first place, such adjustments as those exemplified above are made
comprehensible. Though it is inconceivable that a structure like that of
the pitcher-plant could have been produced by accumulated effects of
function on structure; yet it is conceivable that successive selections
of favourable variations might have produced it; and the like holds of
the no less remarkable appliance of the Venus's Fly-trap, or the still
more astonishing one of that water-plant by which infant-fish are
captured. Though it is impossible to imagine how, by direct influence of
increased use, such dermal appendages as a porcupine's quills could have
been developed; yet, profiting as the members of a species otherwise
defenceless might do by the stiffness of their hairs, rendering them
unpleasant morsels to eat, it is a feasible supposition that from
successive survivals of individuals thus defended in the greatest
degrees, and the consequent growth in successive generations of hairs
into bristles, bristles into spines, spines into quills (for all these
are homologous), this change could have arisen. In like manner, the odd
inflatable bag of the bladder-nosed seal, the curious fishing-rod with
its worm-like appendage carried on the head of the _lophius_ or angler,
the spurs on the wings of certain birds, the weapons of the sword-fish
and saw-fish, the wattles of fowls, and numberless such peculiar
structures, though by no possibility explicable as due to effects of use
or disuse, are explicable as resulting from natural selection operating
in one or other way.
Public-domain text, read in full here on John Shaqi.
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