The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
If we now look to the character of the inhabitants of small
islands{353}, we shall find that those situated close to other land have
a similar fauna with that land{354}, whilst those at a considerable
distance from other land often possess an almost entirely peculiar
fauna. The Galapagos Archipelago{355} is a remarkable instance of this
latter fact; here almost every bird, its one mammifer, its reptiles,
land and sea shells, and even fish, are almost all peculiar and distinct
species, not found in any other quarter of the world: so are the
majority of its plants. But although situated at the distance of between
500 and 600 miles from the S. American coast, it is impossible to even
glance at a large part of its fauna, especially at the birds, without at
once seeing that they belong to the American type{356}. Hence, in fact,
groups of islands thus circumstanced form merely small but well-defined
sub-divisions of the larger geographical divisions. But the fact is in
such cases far more striking: for taking the Galapagos Archipelago as an
instance; in the first place we must feel convinced, seeing that every
island is wholly volcanic and bristles with craters, that in a
geological sense the whole is of recent origin comparatively with a
continent; and as the species are nearly all peculiar, we must conclude
that they have in the same sense recently been produced on this very
spot; and although in the nature of the soil, and in a lesser degree in
the climate, there is a wide difference with the nearer part of the S.
American coast, we see that the inhabitants have been formed on the same
closely allied type. On the other hand, these islands, as far as their
physical conditions are concerned, resemble closely the Cape de Verde
volcanic group, and yet how wholly unlike are the productions of these
two archipelagoes. The Cape de Verde{357} group, to which may be added
the Canary Islands, are allied in their inhabitants (of which many are
peculiar species) to the coast of Africa and southern Europe, in
precisely the same manner as the Galapagos Archipelago is allied to
America. We here clearly see that mere geographical proximity affects,
more than any relation of adaptation, the character of species. How many
islands in the Pacific exist far more like in their physical conditions
to Juan Fernandez than this island is to the coast of Chile, distant 300
miles; why then, except from mere proximity, should this island alone be
tenanted by two very peculiar species of humming-birds--that form of
birds which is so exclusively American? Innumerable other similar cases
might be adduced.
{353} For the general problem of Oceanic Islands, see _Origin_, Ed.
i. p. 388, vi. p. 541.
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