The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
Abortive organs, which must be considered as useless as far as their
ordinary and normal purpose is concerned, are sometimes adapted to other
ends{492}: thus the marsupial bones, which properly serve to support the
young in the mother's pouch, are present in the male and serve as the
fulcrum for muscles connected only with male functions: in the male of
the marigold flower the pistil is abortive for its proper end of being
impregnated, but serves to sweep the pollen out of the anthers{493}
ready to be borne by insects to the perfect pistils in the other
florets. It is likely in many cases, yet unknown to us, that abortive
organs perform some useful function; but in other cases, for instance in
that of teeth embedded in the solid jaw-bone, or of mere knobs, the
rudiments of stamens and pistils, the boldest imagination will hardly
venture to ascribe to them any function. Abortive parts, even when
wholly useless to the individual species, are of great signification in
the system of nature; for they are often found to be of very high
importance in a natural classification{494}; thus the presence and
position of entire abortive flowers, in the grasses, cannot be
overlooked in attempting to arrange them according to their true
affinities. This corroborates a statement in a previous chapter, viz.
that the physiological importance of a part is no index of its
importance in classification. Finally, abortive organs often are only
developed, proportionally with other parts, in the embryonic or young
state of each species{495}; this again, especially considering the
classificatory importance of abortive organs, is evidently part of the
law (stated in the last chapter) that the higher affinities of organisms
are often best seen in the stages towards maturity, through which the
embryo passes. On the ordinary view of individual creations, I think
that scarcely any class of facts in natural history are more wonderful
or less capable of receiving explanation.
{492} The case of rudimentary organs adapted to new purposes is
discussed in the _Origin_, Ed. i. p. 451, vi. p. 620.
{493} This is here stated on the authority of Sprengel; see also
_Origin_, Ed. i. p. 452, vi. p. 621.
{494} _Origin_, Ed. i. p. 455, vi. p. 627. In the margin R. Brown's
name is given apparently as the authority for the fact.
{495} _Origin_, Ed. i. p. 455, vi. p. 626.
_The abortive organs of physiologists._
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