The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
Physiologists and medical men apply the term "abortive" in a somewhat
different sense from naturalists; and their application is probably the
primary one; namely, to parts, which from accident or disease before
birth are not developed or do not grow{496}: thus, when a young animal
is born with a little stump in the place of a finger or of the whole
extremity, or with a little button instead of a head, or with a mere
bead of bony matter instead of a tooth, or with a stump instead of a
tail, these parts are said to be aborted. Naturalists on the other hand,
as we have seen, apply this term to parts not stunted during the growth
of the embryo, but which are as regularly produced in successive
generations as any other most essential parts of the structure of the
individual: naturalists, therefore, use this term in a metaphorical
sense. These two classes of facts, however, blend into each other{497};
by parts accidentally aborted, during the embryonic life of one
individual, becoming hereditary in the succeeding generations: thus a
cat or dog, born with a stump instead of a tail, tends to transmit
stumps to their offspring; and so it is with stumps representing the
extremities; and so again with flowers, with defective and rudimentary
parts, which are annually produced in new flower-buds and even in
successive seedlings. The strong hereditary tendency to reproduce every
either congenital or slowly acquired structure, whether useful or
injurious to the individual, has been shown in the first part; so that
we need feel no surprise at these truly abortive parts becoming
hereditary. A curious instance of the force of hereditariness is
sometimes seen in two little loose hanging horns, quite useless as far
as the function of a horn is concerned, which are produced in hornless
races of our domestic cattle{498}. Now I believe no real distinction can
be drawn between a stump representing a tail or a horn or the
extremities; or a short shrivelled stamen without any pollen; or a
dimple in a petal representing a nectary, when such rudiments are
regularly reproduced in a race or family, and the true abortive organs
of naturalists. And if we had reason to believe (which I think we have
not) that all abortive organs had been at some period _suddenly_
produced during the embryonic life of an individual, and afterwards
become inherited, we should at once have a simple explanation of the
origin of abortive and rudimentary organs{499}. In the same manner as
during changes of pronunciation certain letters in a word may become
useless{500} in pronouncing it, but yet may aid us in searching for its
derivation, so we can see that rudimentary organs, no longer useful to
the individual, may be of high importance in ascertaining its descent,
that is, its true classification in the natural system.
{496} _Origin_, Ed. i. p. 454, vi. p. 625.
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