The Fundamentals of BacteriologyMorrey, Charles Bradfield
Science
The Fundamentals of Bacteriology
Morrey, Charles Bradfield
Bacteriology
=Capsule.=--The _capsule_ is a second covering outside the cell wall
and probably developed from it (Fig. 19). It is usually gelatinous,
so that bacteria which form capsules frequently stick together
when growing in a fluid, so that the whole mass has a jelly-like
consistency. The term _zoöglœa_ was formerly applied to such masses,
but it is a poor term and misleading (zoön = an animal) and should
be dropped. The masses of jelly-like material frequently found on
decaying wood, especially in rainy weather, are in some cases masses
of capsule-forming bacteria, though a part of the jelly is a product
of bacterial activity, a gum-like substance which lies among the
capsulated organisms. When these masses dry out, they become tough
and leathery, but it is not to be presumed that capsules are of this
consistency. On the contrary, they are soft and delicate, though they
certainly serve as an additional protection to the organism, doubtless
more by selective absorption than mechanically. Certain bacteria
which cause disease form capsules in the blood of those animals which
they kill and not in the blood of those in which they have no effect
(_Bacterium anthracis_ in guinea pig’s blood and in rat’s blood). The
presence of capsules around an organism can be proved only by staining
the capsule. Many bacteria when stained in albuminous fluids show a
clear space around them which appears like a capsule. It is due to the
contraction of the fluid away from the organism during drying.
=Metachromatic Granules.=--The term “_metachromatic_” is applied to
granules which in stained preparations take a color different from
the protoplasm as a whole (Fig. 20). They vary widely in chemical
composition. Some of them are glycogen, some fat droplets. Others are
so-called “granulose” closely related to starch but probably not true
starch. Others are probably nuclein. Of many the chemical composition
is unknown. They are called “Babes-Ernst corpuscles” in certain
bacteria (typhoid bacillus). Since they frequently occur in the ends
of cells the term “polar granules” is also applied. Their presence is
of value in the recognition of but few bacteria (“Neisser granules” in
diphtheria).
=Flagellum.=--A _flagellum_ is a very minute thread-like process
growing out from the cell wall, probably filled with a strand of
protoplasm. The vibrations of the flagella move the organism through
the liquid medium. Bacteria which are thus capable of independent
movement are spoken of as “motile bacteria.” The actual rate of
movement is very slight, though in proportion to the size of the
organism it may be considered rapid. Thus Alfred Fischer determined
that some organisms have a speed for short periods of about 40 cm. per
hour. This is equivalent to a man moving more than 200 miles in the
same time.
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