As to specific determinations, a little experience in the field
discloses an amount of variation in species that does not always appear
in the descriptions of authors; and species that are under the closest
scrutiny of botanists, foresters or horticulturalists, attest by their
multiple synonymy their wide variation. The possibilities of variation
are indefinite and, with adaptable Pines, the range of variation is
somewhat proportionate to change of climate. In mountainous countries,
where there are warm sheltered valleys with rich soil below cold barren
ledges, the most variable Pines are found. The western species of North
America, for instance, are much more variable than the eastern species,
while in Mexico, a tropical country with snow-capped mountains, the
variation is greatest.
Therefore in the limitation of species undue importance should not be
given to characters responsive to environment, such as the dimensions of
leaf or cone, the number of leaves in the fascicle, etc. Moreover, there
are familiar examples (P. sylvestris, etc.) that show the possibility of
wide differences in the cone of the same species.
In the following classification species only are considered without
attempting to determine varietal or other subspecific forms. But
varieties are often mentioned as one of the factors illustrating the
scope of species. Synonymy serves a like purpose, but synonyms not
conveying useful information are omitted, Roezl's list of Mexican
species, for instance, and variations in the orthography of specific
names.
PINUS
1755 Pinus Duhamel, Traité des Arbres, ii. 121.
1790 Apinus Necker, Elem. Bot. iii. 269.
1852 Cembra Opiz, Seznam, 27.
1854 Strobus Opiz, Lotos, iv. 94.
1903 Caryopitys Small, Fl. Southeast. U. S. 29.
Leaves and shoots dimorphous, primary leaves on long shoots, secondary
leaves on dwarf shoots. Flowers diclinous, the pistillate taking the
place of long shoots, the staminate taking the place of dwarf shoots.
Growth of wood and fruit emanating from the nodes; buds, branchlets and
cones, therefore, in verticillate association. Leaves and staminate
flowers in internodal position, the primary leaves along the whole
length of the internode, subtending secondary leaf-fascicles on the
apical, staminate flowers on the basal part. Buds compounded of minute
buds in the axils of bud-scales, becoming the bracts of the
spring-shoot. Branchlets of one or more internodes, each internode in
three parts--a length without leaves, a length bearing leaves and a node
of buds. Cone requiring two, rarely three years to mature, displaying
its annual growths by distinct areas on each scale. Seeds wingless or
winged, edible and nutritious.
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