The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
_General Considerations on the Extinct Mammalian Fauna of Europe._--It is a
curious fact that no family, and hardly a genus, of European mammalia
occurs in the Pliocene deposits, without extending back also into those of
Miocene age. There are, {127}however, a few groups which, seem to be late
developments or recent importations into the Palæarctic region, as they
occur only in Post-Pliocene deposits. The most important of these are the
badger, glutton, elk, reindeer, chamois, goat, and sheep, which only occur
in caves and other deposits of Post-Pliocene age. Camels only occur in the
Post-Pliocene of Siberia (_Merycotherium_), although a true _Camelus_ of
large size appears to have inhabited some part of Central Asia in the Upper
Miocene period, being found in the Siwalik beds. The only exclusively
Pliocene genera in Europe are _Ursus_, _Equus_, _Hippopotamus_, _Bos_,
_Elephas_, _Arvicola_, _Trogontherium_, _Arctomys_, _Hystrix_ and _Lepus_;
but of these _Equus_, _Hippopotamus_, _Bos_, and _Elephas_ are found in the
Miocene deposits of India. Owing, no doubt, in part to the superior
productiveness of the various Miocene beds, large numbers of groups appear
to have their origin or earliest appearance here. Such are Insectivora,
Felidæ, Hyænidæ, Mustelidæ, _Ursus_, Equidæ, _Tapirus_, Rhinocerotidæ,
Hippopotamidæ, Anthracotheridæ (extinct), _Sus_, Camelopardidæ, Tragulidæ,
Cervidæ, Bovidæ, Elephantidæ, and Edentata.
Groups which go back to the Eocene period, are, Primates allied to South
American monkeys, as well as some of the Lemuridæ; bats of the living genus
_Vespertilio_; Hyænodontidæ, an ancestral form of Carnivore; Viverridæ;
Canidæ (to the Upper Eocene), and the ancestral Arctocyonidæ to the Lower
Eocene; _Hyænarctos_, an ancestral type of bears and hyænas; Anchitheridæ,
ancestral horses, to the Middle Eocene; Palæotheridæ, comprising numerous
generalised forms, ancestors of the rhinoceros, horse, and tapir; Suidæ,
with numerous generalised forms, to the Middle Eocene; Anoplotheridæ and
Xiphodontidæ, ancestral families of even-toed Ungulates, connecting the
ruminants with the swine; and lastly, several groups of Rodents, and a
Marsupial, in the Upper Eocene. We thus find all the great types of
Mammalia well developed in the earliest portion of the tertiary period; and
the occurrence of Quadrumana, of the highly specialized bats
(_Vespertilio_), of various forms of Carnivora, and of Ungulates, clearly
differentiated into the odd and even-toed series, associated with such
lower forms as {128}Lemurs and Marsupials--proves, that we have here hardly
made an approach towards the epoch when the mammalian type itself began to
diverge into its various modifications. Some of the Carnivora and Ungulates
do, indeed, exhibit a less specialised structure than later forms; yet so
far back as the Upper Miocene the most specialised of all carnivora, the
great sabre-toothed _Machairodus_, makes its appearance.
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