The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
Allied to these, are a series of forms showing a transition to the tapirs,
and to the _Palæotherium_ of the European Eocene. In the Pliocene we have
_Parahippus_; in the Miocene _Lophiodon_, found in the same formation and
in the Eocene of Europe, and allied to the tapir; and in the Eocene,
_Palæosyops_, as large as a rhinoceros, which had large canines and was
allied to the tapir and _Palæotherium_; _Limnohyus_, forming the type of a
family Limnohyidæ, which included the last genus and some others mentioned
further on; and _Hyrachyus_, allied to _Lophiodon_, and to _Hyracodon_ an
extinct form of rhinoceros. Besides these we have _Lophiotherium_ (also
from the Eocene of Europe); _Diplacodon_ allied to _Limnohyus_, but with
affinities to modern Perissodactyla and nearly as large as a rhinoceros;
and _Colonoceras_, also belonging to the Limnohyidæ, an animal which was
the size of a sheep, and had divergent protuberances or horns on its nose.
A remarkable genus, _Bathmodon_, lately described by Professor Cope, and of
which five species have been found in the Eocene of New Mexico and Wyoming,
is believed to form the type of a new family, having some affinity to
_Palæosyops_ and to the extinct Brontotheridæ. It had large canine tusks
but no horns.
The Rhinocerotidæ are represented in America by the genus _Rhinoceros_ in
the Pliocene and Miocene, and by _Aceratherium_ and _Hyracodon_ in the
Miocene. Both the latter were hornless, and _Hyracodon_ was allied to the
Eocene _Hyrachyus_, one of the Lophiodontidæ. In the Eocene and Miocene
deposits of Utah, and Oregon, several remarkable extinct rhinoceroses have
been {137}recently discovered, forming the genus _Diceratherium_. These had
a pair of nasal horns placed side by side on the snout, not behind each
other as in existing two-horned rhinoceroses, the rest of their skeleton
resembling the hornless _Aceratherium_. They were of rather small size.
Next to these extinct rhinoceroses come the Brontotheridæ, an extraordinary
family of large mammalia, some of which exceeded in bulk the largest living
rhinoceros. They had four toes to the front and three to the hind feet,
with a pair of large divergent horns on the front of the head, in both
sexes. Professor Marsh and Dr. Leidy have described four genera,
_Brontotherium_, _Titanotheium_, _Megacerops_, and _Anisacodon_,
distinguished by peculiarities of dentition. Though most nearly allied to
the rhinoceroses, they show some affinity for the gigantic Dinocerata of
the Eocene to be noticed further on. Professor Cope has since described
another genus, _Symborodon_, from the Miocene of Colorado, with no less
than seven species, one nearly the size of an elephant. He thinks they had
a short tapir-like proboscis. The species differ greatly in the form of the
cranium and development of the horn-bearing processes.
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