The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
_Proboscidea._--The Elephantidæ are only represented in America by one
species of _Mastodon_ and one of _Elephas_, in the Newer Pliocene deposits.
In the Older Pliocene, Miocene, {139}and Upper Eocene, no remains of this
order have been found; and in 1869, Dr. Leidy remarked on the small average
size of the extinct North American mammalia, which were almost all smaller
than their living analogues. Since then, however, wonderful discoveries
have been made in deposits of Middle Eocene age in Wyoming and Colorado, of
a group of huge animals not only rivalling the elephants in size, but of so
remarkable and peculiar a structure as to require the formation of a new
order of mammals--Dinocerata--for their reception.
This order consists of animals with generalised Ungulate and Proboscidean
affinities. The lower jaw resembles that of the hippopotamus; they had five
toes on the anterior feet and four on the posterior; three pairs of horns,
the first pair on the top of the head, large and perhaps palmated, the
second pair above the eyes, while the third and smallest stood out sideways
on the snout. They had enormous upper canines, of which the roots entered
the middle horn cores, no upper incisors, and small molars. Professor Marsh
believes that they had no trunk. The remains discovered indicate four
genera, _Dinoceras_ (3 sp.), _Tinoceras_ (2 sp.), _Uintatherium_ (1 sp.),
and _Eobasileus_ (2 sp.). Many other names have been given to fragments of
these animals, and even those here given may not be all distinct.
Another new order, Tillodontia, recently established by Professor Marsh, is
perhaps yet more remarkable in a zoological point of view, since it
combines the characters of Carnivora, Ungulata, and Rodents. These animals
have been formed into two families, Tillotheridæ and Stylinodontidæ; and
three genera, _Tillotherium_, _Anchippodus_, and _Stylinodontia_. All are
from the Eocene of Wyoming and New Jersey. Perhaps to these must be added
_Elotherium_ from the Miocene of Dakota, the other forms being all Eocene.
They were mostly animals of small size, between that of the capybara and
tapir. The skull resembled in form that of a bear; the molar teeth were of
Ungulate type, and the incisors like those of a Rodent; but the skeleton
was more that of the Ursidæ, the feet being plantigrade. Professor Cope has
since described three new genera from the Eocene of New Mexico,
_Ectoganus_, _Calamodon_, and _Esthonyx_, comprising {140}seven species
allied to _Tillotherium_ and _Anchippodus_, and having also relations, as
Professor Cope believes, with the South American Toxodontidæ.
Public-domain text, read in full here on John Shaqi.
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