The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
_Coleoptera of St. Helena._--Omitting those beetles which get introduced
everywhere through man's agency, there are 59 species of Coleoptera known
from St. Helena; and even of these there are a few widely distributed
species that may have been introduced by man. It will be well, therefore,
to confine ourselves almost wholly to the species peculiar to the island,
and, therefore, almost certainly forming part of the endemic or original
fauna. Of these we find that 10 belong to genera which have a very wide
range, and thus afford no indication of geographical affinity; 2 belong to
genera which are characteristic of the Palæarctic fauna (_Bembidium_,
_Longitarsus_); 3 to African genera (_Adoretus_, _Sciobius_,
_Aspidomorpha_); and two species of _Calosoma_ are most allied to African
species. There are also 4 African species, which may be indigenous in St.
Helena. The peculiar genera, 7 in number, are, however, the most
interesting. We have first _Haplothorax_, a large beetle allied to
_Carabus_ and _Calosoma_, though of a peculiar type. This may be held to
indicate a remote Palæarctic affinity. _Melissius_, one of the Dynastidæ,
is allied to South African forms. _Microxylobius_, one of the Cossonides (a
sub-family of Curculionidæ) is the most important genus, comprising as it
does 13 species. It is, according to Mr. Wollaston, an altogether peculiar
type, most allied to _Pentarthrum_, a genus found in St. Helena, Ascension,
and the south of England, and itself very isolated. _Nesiotes_, another
genus of Curculionidæ, belongs to a small group, the allied genera forming
which inhabit Europe, Madeira, and Australia. A third peculiar and isolated
genus is _Trachyphlæosoma_. The Anthribidæ are represented by {271}2
genera, _Notioxenus_ and _Homoeodera_, which are altogether peculiar and
isolated, and contain 9 species. Thus no less than 27 species, or more than
half of the undoubtedly indigenous beetles, belong to 5 peculiar and very
remarkable genera of Rhyncophora.
It appears from this enumeration, that the peculiar species as a whole,
exhibit most affinity to the Ethiopian fauna; next to the South European
fauna; and lastly to that of the islands of the North Atlantic; while there
is such a large amount of peculiarity in the most characteristic forms,
that no special geographical affinity can be pointed out.
_Land Shells._--These consist of about a dozen living species, and about as
many extinct found in the surface soil, and probably exterminated by the
destruction of the forests. The genera are _Succinea_, _Zonites_, _Helix_,
_Bulimus_, _Pupa_, and _Achatina_. The _Bulimi_ (all now extinct but one)
comprise one large, and several small species, of a peculiar type, most
resembling forms now inhabiting South America and the islands of the
Pacific. _Zonites_ is chiefly South European, but the other genera are of
wide range, and none are peculiar to the island.
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