The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
fresh water molluscs and their eggs must frequently be carried out to sea,
yet this cannot lead to their dispersal, since salt water is almost
immediately fatal to them; and we are therefore forced to conclude that the
apparently insignificant and uncertain means of dispersal above alluded to
are really what have led to their wide distribution. The true land-shells
offer a still more difficult case, for they are exceedingly sensitive to
the influence of salt water; they are not likely to be carried by aquatic
birds, and yet they are more or less abundant all over the globe,
inhabiting the most remote oceanic islands. It has been found, however,
that land-shells have the power of lying dormant a long time. Some have
lived two years and a half shut up in pill boxes; and one Egyptian desert
snail came to life after having been glued down to a tablet in the British
Museum for four years!
We are indebted to Mr. Darwin for experiments on the power of land shells
to resist sea water, and he found that when they had formed a membranous
diaphragm over the mouth of the shell they survived many days' immersion
(in one case fourteen days); and another experimenter, quoted by Mr.
Darwin, found that out of one hundred land shells immersed for a fortnight
in the sea, twenty-seven recovered. It is therefore quite possible for them
to be carried in the chinks of drift wood for many hundred miles across the
sea, and this is probably one of the most effectual modes of their
dispersal. Very young shells would also {32}sometimes attach themselves to
the feet of birds walking or resting on the ground, and as many of the
waders often go far inland, this may have been one of the methods of
distributing species of land shells; for it must always be remembered that
nature can afford to wait, and that if but once in a thousand years a
single bird should convey two or three minute snails to a distant island,
this is all that is required for us to find that island well stocked with a
great and varied population of land shells.
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