The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
These are large angled nuts contained in a thick envelope, and showing
internal structures resembling those of the seeds of modern _Taxineæ_.
There are numerous species, as well as allied seeds referred to the
provisional genera _Rhabdocarpus_ and _Carpolithes_. In _Trigonocarpum
Hookeri_ I have described the internal structure of one of those seeds,
and many fine examples from the coal-field of St. Etienne, in France,
have been described by Brongniart, so that their internal structure is
very well known.
_Genus_ Antholithes.
This is also a provisional genus, to include spikes of floral organs,
some of which are known to have belonged to _Cordaites_, others probably
to _Sigillariæ_.
Of Uncertain Affinities.
_Family_ Sigillariaceæ.
Under this name palæobotanists have included a great number of trees
of the Carboniferous system, all of which are characterised by broad
leaf-sears, with three vascular scars, and usually arranged in
vertical rows, and by elongated three-nerved leaves, and roots of the
stigmaria type--that is, with rounded pits, marking the attachment of
rootlets spirally arranged. These trees, however, collected in the
genus Sigillaria by arbitrary characters, which pass into those of
the Lepidodendroid trees, have been involved in almost inextricable
confusion, to disentangle which it will be necessary to consider: 1. The
external characters of _Sigillariæ_, and trees confounded with them. 2.
Subdivision of _Sigillariæ_ by external markings. 3. The microscopic
character of their stems. 4. What is known of their foliage and fruit.
1. _Characters of Sigillaroid and Lepidodendroid Trunks_.
It may be premised that the modes of determination in fossil botany
are necessarily different from those employed in recent botany. The
palæobotanist must have recourse to characters derived from the leaves,
the scars left by their fall, and the internal structures of the stem.
These parts, held in little esteem by botanists in describing modern
plants, and much neglected by them, must hold the first place in the
regard of the fossil botanist, whereas the fructification, seldom
preserved, and generally obscure, is of comparatively little service. It
is to be remarked also that in such generalised plants as those of the
Palæozoic, remarkable rather for the development of the vegetative than
of the reproductive organs, the former rise in importance as compared
with their value in the study of modern plants.
In _Sigillariæ_, _Lepidodendra_, &c., the following surfaces of the stem
may be presented to our inspection:
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