The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
1. The outer surface of the epidermis without its leaves, but with
the leaf-bases and leaf-scars more or less perfectly preserved. On
this surface we may recognise: (1) Cellular swellings or projections
of the bark to which the leaves are attached. These may be called
leaf-bases, and they are sometimes very prominent. (2) The actual mark
of the attachment of the leaf situated in the most prominent part of
the leaf-base. This is the _leaf-scar_. (3) In the leaf-scar when well
preserved we can see one or more minute punctures or prominences which
are the points where the vascular bundles passing to the leaf found exit.
These are the vascular scars.
When the leaves are attached, the leaf-scars and vascular scars cannot
be seen, but the leaf-bases can be made out. Hence it is important, if
possible, to secure specimens with and without the leaves. In flattened
specimens the leaf-bases are often distorted by pressure and marked with
furrows which must not be mistaken for true structural characters. The
leaf-bases, which are in relief on the outer surface of the stem, of
course appear as depressions on the mould in the containing rock, in
which the markings often appear much more distinctly than on the plant
itself.
2. The outer surface of the epidermis may have been removed or may be
destroyed by the coarseness of the containing rock. In this case the
leaf-bases are usually preserved on the surface of the outer or corky
bark, but the leaf-scars and vascular scars have disappeared. This gives
that condition of Lepidodendroid trees to which the name _Knorria_ has
been applied. When plants are in this state careful inspection may
sometimes discover traces of the leaf-scars on portions of the stem, and
thus enable the _Knorria_ to be connected with the species to which it
belongs.
3. The outer or corky bark may be removed, exposing the surface of the
inner or fibrous and cellular bark, which in the plants in question is
usually of great thickness. In this case neither the leaf-bases nor the
scars are seen, but punctures or little furrows or ridges appear where
the vascular bundles entered the inner bark. Specimens in this state are
usually said to be decorticated, though only the outer bark is removed.
It is often difficult to determine plants in this condition, unless some
portion of the stem can be found still retaining the bark; but when care
is taken in collecting, it will not infrequently be found that the true
outer surface can be recovered from the containing rock, especially if
a coaly layer representing the outer bark intervenes between this and
the inner impression. Specimens of this kind, taken alone, have been
referred to the genera _Knorria_, _Bothrodendron_, and _Halonia_.
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