The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
4. In some cases, though not frequently, the outer surface of the
ligneous cylinder is preserved. It almost invariably presents a regularly
striated or irregularly wrinkled appearance, depending upon the vertical
woody wedges, or the positions of the medullary rays or vascular bundles.
Specimens of this kind constituted some of the _Endogenites_ of the
older botanists, and the genus _Schizodendron_ of Eichwald appears to
include some of them. Many of them have also been incorrectly referred to
Calamites.
5. In some cases the cast of the medullary cylinder or pith may alone
be preserved. This may be nearly smooch or slightly marked by vertical
striæ, but more usually presents a transverse striation, and not
infrequently the transverse constrictions and septa characteristic of the
genus Sternbergia. Loose _Sternbergiæ_ afford little means of connecting
them with the species to which they belong, except by the microscopic
examination of the shreds of the ligneous cylinder which often cling to
them.[CS]
[CS] See my paper, "Journal of Geological Society," vol. xxvii.
These facts being premised, the following general statements may be made
respecting some of the more common Palæozoic genera, referring, however,
principally to the perfect markings as seen on the epidermis:
_Sigillaria._--Leaf-bases hexagonal or elongated, or confluent on a
vertical ridge. Leaf-scars hexagonal or shield-shaped. Vascular scars
three, the two lateral larger than the central. This last character
is constant, depending on the fact that the leaves of Sigillaria have
two or more vascular bundles. All so-called _Sigillariæ_ having the
central vascular scar largest, or only one vascular bundle, should be
rejected from this genus. In young branches of branching _Sigillariæ_ the
leaf-scars sometimes appear to be spiral, but in the older stems they
form vertical rows; interrupted, however, by transverse rows or bands
of _fruit-scars_, each with a single large central vascular scar, and
which have borne the organs of fructification. _Arthrocaulis_ of McCoy is
founded on this peculiarity.
_Syringodendron._--Differs from Sigillaria in the leaf-scars, which are
circular and with a single vascular bundle. It is a matter of doubt
whether these plants were of higher rank than Sigillaria tending toward
the pines, or of lower rank tending toward Cyclostigma. Their leaf-bases
form vertical ridges.
_Lepidodendron._--Leaf-bases rhombic, oval, or lanceolate, moderately
prominent. Leaf-scars rhombic or sometimes shield-shaped or heart-shaped,
in the middle or upper part of the leaf-base. Vascular scars three--the
middle one always largest and corresponding to the single nerve of the
leaf; the lateral ones sometimes obsolete.
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