The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
The genus _Buthotrephis_ of Hall, which is characterised as having
stems, sub-cylindric or compressed, with numerous branches, which are
divaricating and sometimes leaf-like, contains some true Algæ. Hall's
_B. gracilis_, from the Siluro-Cambrian, is one of these. Similar
plants, referred to the same species, occur in the Clinton and Niagara
formations, and a beautiful species, collected by Col. Grant, of
Hamilton, and now in the McGill College collection, represents a broader
and more frondose type of distinctly carbonaceous character. It may be
described as follows:
_Buthotrephis Grantii_, S. N. (Fig. 13).--Stems and fronds smooth and
slightly striate longitudinally, with curved and interrupted striæ.
Stem thick, bifurcating, the divisions terminating in irregularly
pinnate fronds, apparently truncate at the extremities. The quantity
of carbonaceous matter present would indicate thick, though perhaps
flattened, stems and dense fleshy fronds.
[Illustration: Fig. 13.--_Buthotrephis Grantii_, a genuine Alga from the
Silurian, Canada.]
The species _Buthotrephis subnodosa_ and _B. flexuosa_, from the Utica
shale, are also certainly plants, though it is possible, if their
structures and fruit were known, some of these might be referred to
different genera. All of these plants have either carbonaceous matter or
produce organic stains on the matrix.
The organism with diverging wedge-shaped fronds, described by Hall as
_Sphenothallus angustifolius_, is also a plant. Fine specimens, in the
collection of the Geological Survey of Canada, show distinct evidence of
the organic character of the wedge-shaped fronds. It is from the Utica
shale, and elsewhere in the Siluro-Cambrian. It is just possible, as
suggested by Hall, that this plant may be of higher rank than the Algæ.
The genus _Palæophycus_ of Hall includes a great variety of uncertain
objects, of which only a few are probably true Algæ. I have specimens of
fragments similar to his _P. virgatus_, which show distinct carbonaceous
films, and others from the Quebec group, which seem to be cylindrical
tubes now flattened, and which have contained spindle-shaped sporangia
of large size. Tortuous and curved flattened stems, or fronds, from the
Upper Silurian limestone of Gaspé, also show organic matter.
Respecting the forms referred to _Licrophycus_ by Billings, containing
stems or semi-cylindrical markings springing from a common base, I have
been in great doubt. I have not seen any specimens containing unequivocal
organic matter, and am inclined to think that most of them, if not the
whole, are casts of worm-burrows, with trails radiating from them.
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