The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
The history of the ferns in geological time is remarkably different
from that of the Lycopods; for while the latter have long ago descended
from their pristine eminence to a very humble place in nature, the
former still, in the southern hemisphere at least, retain their arboreal
dimensions and ancient dominance.
[Illustration: Fig. 28.--Asterophyllites (Erian, New Brunswick), A,
Asterophyllites latifolia. B, Do., apex of stem (?) fruit, C,
C^1, _A. scutigera_. D, _A. latifolia_, larger whorl of leaves. D^1,
Leaf.]
The family of the _Equisetaceæ_, or mare's-tails, was also represented
by large species of _Calamites_ and by _Asterophyllites_ in the Erian;
but, as its headquarters are in the Carboniferous, we may defer its
consideration till the next chapter. (Figs. 27, 28.)
Passing over these for the present, we find that the flowering plants are
represented in the Erian forests by at least two types of Gymnosperms,
that of _Taxineæ_ or yews, and an extinct family, that of the _Cordaites_
(Figs. 30, 31). The yew-trees are closely allied to the pines and
spruces, and are often included with them in the family of _Coniferæ_.
They differ, however, in the habit of producing berries or drupe-like
fruits instead of cones, and there is some reason to believe that this
was the habit of the Erian trees of this group, though their wood in
some instances resembles rather that of the Araucaria, or Norfolk
Island pine, than that of the modern yews. These trees are chiefly
known to us by their mineralised trunks, which are often found like
drift-wood on modern sand-banks embedded in the Erian sandstones or
limestones. It often shows its structure in the most perfect manner in
specimens penetrated by calcite or silica, or by pyrite, and in which
the original woody matter has been resolved into anthracite or even into
graphite. These trees have true woody tissues presenting that beautiful
arrangement of pores or thin parts enclosed in cup-like discs, which is
characteristic of the coniferous trees, and which is a great improvement
on the barred tissue already referred to, affording a far more strong,
tough, and durable wood, such as we have in our modern pines and yews
(Fig. 29).
[Illustration: Fig. 29.--_Dadoxylon Ouangondianum_, an Erian conifer, A,
Fragment showing Sternberg pith and wood; _a_, medullary sheath; _b_,
pith; _c_, wood; _d_, section of pith, B, Wood-cell; _a_, hexagonal
areole; _b_, pore, _c_, Longitudinal section of wood, showing, _a_,
areolation, and _b_, medullary rays, D, Transverse section, showing,
_a_, wood-cells, and _b_, limit of layer of growth, (B, C, D, highly
magnified.)]
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