The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
These primitive pines make their appearance in the Middle Erian, in
various parts of America, as well as in Scotland and Germany, and they
are represented by wood indicating the presence of several species. I
have myself indicated and described five species from the Erian of Canada
and the United States. From the fact that these trees are represented
by drifted trunks embedded in sandstones and marine limestones, we may,
perhaps, infer that they grew on the rising grounds of the Erian land,
and that their trunks were carried by river-floods into the sea. No
instance has yet certainly occurred of the discovery of their foliage or
fruit, though there are some fan-shaped leaves usually regarded as ferns
which may have belonged to such trees. These in that case would have
resembled the modern _Gingko_ of China, and some of the fruits referred
to the genus _Cardiocarpum_ may have been produced by them. Various
names have been given to these trees. I have preferred that given by
Unger, _Dadoxylon_, as being more non-committal as to affinities than the
others.[BC] Many of these trees had very long internal pith-cylinders,
with curious transverse tubulæ, and which, when preserved separately,
have been named _Sternbergia_.
[BC] _Araucarites_, Goeppert; _Araucarioxylon_, Kraus.
Allied to these trees, and perhaps intermediate between them and the
_Cycads_, were those known as _Cordaites_ (Fig. 30), which had trunks
resembling those of _Dadoxylon_, but with still larger _Sternbergia_
piths and an internal axis of scalariform vessels, surrounded by a
comparatively thin woody cylinder. Some of them have leaves over a
foot in length, reminding one of the leaves of broad-leaved grasses or
iridaceous plants. Yet their flowers and fruit seem to have been more
nearly allied to the yews than to any other plants (Fig. 31). Their stems
were less woody and their piths larger than in the true pines, and some
of the larger-leaved species must have had thick, stiff branches. They
are regarded as constituting a separate family, intermediate between
pines and cycads, and, beginning in the Middle Devonian, they terminate
in the Permian, where, however, some of the most gigantic species occur.
In so far as the form and structure of the leaves, stems, and fruit are
concerned, there is marvellously little difference between the species
found in the Erian and the Permian. They culminated, however, in the
Carboniferous period, and the coal-fields of southern France have proved
so far the richest in their remains.
[Illustration: Fig. 30.--_Cordaites Robbii_ (Erian, New Brunswick),
_a_, Group of young leaves. _b_, Point of leaf, _c_, Base of leaf, _d_.
Venation, magnified.]
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