The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
Prominent among the more gigantic trees of these swampy forests are
those known to us as _Sigillariæ_ (Fig. 33). They have tall, pillar-like
trunks, often several feet in diameter, ribbed like fluted columns, but
in the reverse way, and spreading at the top into a few thick branches,
which are clothed with long, grass-like leaves. They resemble in some
respects the Lepidodendra of the Erian age, but are more massive, with
ribbed instead of scaly trunks, and longer leaves. If we approach one
of them more closely, we are struck with the regular ribs of its trunk,
dotted with rows of scars of fallen leaves, from which it receives its
name _Sigillaria_, or seal-tree (Figs. 34-37). If we cut into its stem,
we find that, instead of the thin bark and firm wood with which we are
familiar in our modern trees, it has a hard external rind, then a great
thickness of cellular matter with rope-like bands of fibres, constituting
an inner bark, while in the centre is a firm, woody axis of comparatively
small diameter, and somewhat intermediate in its structures between that
of the Lepidodendra and those of the cycads and the taxine conifers.
Thus a great stem, five feet in diameter, may consist principally of
cellular and bast fibres with very little true woody matter. The roots
of this tree are perhaps its most singular feature. They usually start
from the stem in four main branches, then regularly bifurcate several
times, and then run out into great cylindrical cables, running for a
long distance, and evidently intended to anchor the plant firmly in a
soft and oozy soil. They were furnished with long, cylindrical rootlets
placed regularly in a spiral manner, and so articulated that when they
dropped off they left regular rounded scars. They are, in short, the
_Stigmariæ_, which we have already met with in the Erian (Figs. 38, 39).
In Fig. 33 I have endeavoured to restore these strange trees. It is not
wonderful that such plants have caused much botanical controversy. It was
long before botanists could be convinced that their roots are properly
roots at all, and not stems of some aquatic plant. Then the structure of
their stems is most puzzling, and their fruit is an enigma, for while
some have found connected with them cones supposed to resemble those of
lycopods, others attribute to them fruits like those of yew-trees. For
years I have been myself gathering materials from the rich coal-formation
deposits of Nova Scotia in aid of the solution of these questions, and
in the mean time Dr. Williamson, of Manchester, and Renault and other
botanists in France, have been amassing and studying stores of specimens,
and it is still uncertain who may finally be the fortunate discoverer
to set all controversies at rest. My present belief is, that the true
solution consists in the fact that there are many kinds of _Sigillariæ_.
While in the modern forests of America and Europe the species of any of
our ordinary trees, as oaks, birches, or maples, may almost be counted
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