The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
on one's fingers, Schimper in his vegetable palæontology enumerates
about eighty species of Carboniferous _Sigillariæ_; and while on the one
hand many of these are so imperfectly known that they may be regarded
as uncertain, on the other hand many species must yet remain to be
discovered.[CA] Now, in so vast a number of species there must be a
great range of organisation, and, indeed, it has already been attempted
to subdivide them into several generic groups. The present state of the
question appears to me to be this, that in these _Sigillariæ_ we have
a group divisible into several forms, some of which will eventually be
classed with the Lepidodendra as lycopods, while others will be found
to be naked-seeded phænogams, allied to the pines and cycads, and to a
remarkable group of trees known as _Cordaites_, which we must shortly
notice.
[CA] In a recent memoir (Berlin, 1887) Stur has raised the number of
species in one subdivision of the _Sigillariæ_ (the _Favulariæ_) to
forty-seven!
[Illustration: Fig. 38.--_Stigmaria_ root, seen from above, showing its
regular divisions. From "Acadian Geology".]
[Illustration: Fig. 39.--Portion of bark of _Stigmaria_, showing scars of
attachment of rootlets.]
Before considering other forms of Carboniferous vegetation, let us
glance at the accumulation of coal, and the agency of the forests of
_Sigillariæ_ therein. Let us imagine, in the first instance, such trees
as those represented in the figures, growing thickly together over vast
swampy flats, with quantities of undergrowth of ferns and other plants
beneath their shade, and accumulating from age to age in a moist soil
and climate a vast thickness of vegetable mould and trunks of trees, and
spores and spore-cases, and we have the conditions necessary for the
growth of coal. Many years ago it was observed by Sir William Logan that
in the coal-field of South Wales it was the rule with rare exceptions
that, under every bed of coal, there is a bed of clay filled with roots
of the _Stigmaria_, already referred to as the root of _Sigillaria_. This
discovery has since been extended to all the coal-fields of Europe and
America, and it is a perfectly conclusive fact as regards the origin of
coal. Each of these "under-clays," as they are called, must, in fact,
have been a soil on which grew, in the first instance, Sigillariæ and
other trees having stigmaria-roots. Thus, the growth of a forest of
_Sigillariæ_ was the first step toward the accumulation of a bed of
coal. More than this, in some of the coarser and more impure coals,
where there has been sufficient earthy matter to separate and preserve
impressions of vegetable forms, we can see that the mass of the coal is
made up of flattened _Sigillariæ_, mixed with vegetable _débris_ of all
kinds, including sometimes vast quantities of lepidodendroid spores,
and the microscopic study of the coal gives similar results (Fig. 40).
Further, on the surfaces of many coals, and penetrating the shales or
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