The last man who has paid special attention to the grapes of North
America is Bailey, of Cornell. In his latest classification he gives
twenty-three species of American grapes. Bailey is the only American
botanist of experience and recognized standing in general botany who has
paid special attention to the grape. His monograph of the genus Vitis
which, with some changes, has appeared successively in _Gray's
Synoptical Flora_, _The Evolution of Our Native Fruits_, and the
_Cyclopedia of American Horticulture_, is the most complete work we have
on this subject. With his permission we have followed his arrangement of
species in _The Grapes of New York_.
With this brief history of the formation of the genus Vitis as it now
stands we pass to a consideration of the botanical characters of Vitis.
From the time when botanists first commenced to work on the grape there
has been a constant search for taxonomic characters for separating the
various species clearly and distinctly. Many of the earlier
descriptions, while they are correct so far as they go, do not mention
enough characters to enable one to distinguish between similar species.
It has been found that dependence upon the shape of the leaf, size of
berry, size of plant, date of ripening, and similar characters, is very
uncertain and unsatisfactory and that, while these characters are always
mentioned in descriptions as indicating the intrinsic value of a
species, they are of little value from a systematic standpoint. There
are, however, several characters of Vitis which have great taxonomic
importance.
One of the fundamental characters which determine a species is
continuous or intermittent tendrils, first noticed by Professor A.
Braun[120] of Berlin. _Vitis labrusca_, the common Fox grape, is
peculiar in that there are tendrils, or an inflorescence, opposite
nearly every leaf; this arrangement is known as _continuous tendrils_.
All other species have two leaves with a tendril opposite each and a
third leaf without a tendril; such species are said to have
_intermittent tendrils_. Continuity of tendrils is a variable character
and to ascertain which of the two forms is present it is necessary to
have vigorous, healthy, typical canes. The lowest leaves of canes
usually have no opposite tendrils. This character is shown in the
color-plates of the several species.
A closely related character is that of the number of inflorescences
borne by a species. All species excepting _Vitis labrusca_ average two
inflorescences to the cane but the last named species, at least in some
of its subdivisions, may bear from three to six inflorescences, each of
course in the place of a tendril opposite a leaf.
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