Attention is called to the characters given by Bartram as distinguishing
the European from the American species. The first difference that he
mentions is in the shape of the fruit, that of the Vinifera being more
or less oval and that of American species roundish or oblate. Recent
technical descriptions of our American species give the fruit as
spherical where the shape is mentioned at all. On the other hand it is
known that most of the cultivated varieties of European grapes are oval.
Does this mean that all of our cultivated American varieties which show
oval berries, such as Isabella, Catawba, and others, contain Vinifera
blood? It could not be said without careful study that this is true but
it is certainly worthy of consideration. This point seems to have
escaped the attention of our later-day botanists.
The sexual status of the grape has always been a source of
misunderstanding. The earlier botanists spoke of American vines as
dioecious, that is, bearing staminate and pistillate flowers on separate
individuals. In this, as was noted on page 98, they were corrected by
Bartram, in so far as American species were concerned, he stating that
the vines of America were polygamous (showing staminate and
hermaphrodite plants). Bartram did not presume to speak as to the sex of
the flowers of the Old World grape. Later it was determined that the
cultivated varieties of Europe were always hermaphrodite and that
staminate forms were unknown. Engelmann[121] explains this so well and
with such apparent satisfaction that we cannot do better than quote him
here. "All the true Grape-vines bear fertile flowers on one stock and
sterile flowers on another separate stock, and are therefore called
_polygamous_, or, not quite correctly, _dioecious_. The sterile plants
do bear male flowers with abortive pistils, so that while they never
produce fruit themselves, they may assist in fertilizing the others; the
fertile flowers, however, are hermaphrodites containing both
organs--stamens and pistils--and are capable of ripening fruit without
the assistance of the male plants. Real female flowers without any
stamens do not seem ever to have been observed. Both forms, the male and
the hermaphrodite, or if preferred those with sterile and those with
complete flowers, are found mixed in their native localities of the wild
plants, but of course only the fertile plants have been selected for
cultivation, and thus it happens that to the cultivator only these are
known; and as the Grape-vine of the Old World has been in cultivation
for thousands of years, it has resulted that this hermaphrodite
character of its flowers has been mistaken for a botanical peculiarity,
by which it was to be distinguished, not only from our American
Grape-vines, but also from the wild grapes of the old world. But plants
raised from the seeds of this as well as any other true Grape-vine,
generally furnish as many sterile as fertile specimens, while those
Public-domain text, read in full here on John Shaqi.
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