The History of the European FaunaScharff, R. F. (Robert Francis)
History
The History of the European Fauna
Scharff, R. F. (Robert Francis)
Animals -- Europe; Zoogeography
Baron Aucapitaine has since tried
similar experiments: he placed one hundred land-shells, belonging to ten
species, in a box pierced with holes, and immersed it for a fortnight
in the sea. Out of the hundred shells, twenty-seven recovered. The
presence of an operculum seems to have been of importance, as out of
twelve specimens of _Cyclostoma elegans_ which it thus furnished, eleven
revived. It is remarkable, seeing how well the _Helix pomatia_ resisted
with me the salt-water, that not one of fifty-four specimens belonging
to four other species of _Helix_ tried by Aucapitaine, recovered. It is,
however, not at all probable that land-shells have often been thus
transported; the feet of birds offer a more probable method."
We have here positive evidence that such shells as _Helix pomatia_ and
_Cyclostoma elegans_ might easily be transported to an island from the
mainland. The former occurs in France, Holland, and England, and the
latter all along western continental Europe and England. And yet neither
of these species inhabits the Canary Islands, Madeira, or Ireland, none
of which are at too great a distance from Europe to be within easy reach
for a floating object. The fact that _Cyclostoma elegans_ does not live
in Ireland is of particular interest in connection with the
floating-theory just quoted, as on all sides of Ireland dead specimens
have been picked up on the shore, showing that marine currents carry
specimens and have thus transported them for countless centuries.
Nevertheless the species has not established itself in Ireland. If such
a fate meets a land-shell of the type of _Cyclostoma elegans_, it may be
asked, with some justification, what chance slugs or the smaller
non-operculated species would have to reach an island like Ireland
alive from the mainland, and to colonise it successfully.
Both slugs and their eggs are killed by a short immersion in sea-water,
as I have proved experimentally. I have also subjected slugs, in the act
of crawling on twigs, to an artificial spray of sea-water. This seemed
to irritate their tender skins to such an extent that they curled
themselves up, released their hold on the twig and let themselves drop
to the ground. If we supposed, therefore, that a slug had successfully
reached the sea, transported on a tree-trunk, the moisture would tend to
lure it forth from its hiding-place under the bark, whilst the mere
spray would prove fatal to its existence. Those species of snails and
slugs which lead an underground existence, rarely venturing above
ground, such as _Testacella_ and _Coecilianella_, would have even less
chance of being accidentally carried to some distant island.
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