The History of the European FaunaScharff, R. F. (Robert Francis)
History
The History of the European Fauna
Scharff, R. F. (Robert Francis)
Animals -- Europe; Zoogeography
We may thus distinguish between the natural or active and the accidental
or passive means of distribution of animals. The active mode of
dispersal again may be only migratory, as in most animals, or periodic
and migratory, as in some birds and fishes. It is of course the tendency
of every species to spread in all directions from its original home,
provided it does not encounter obstacles, such as want of food,
unsuitability of climate or soil, or barriers such as mountains, rivers,
or the sea. Birds might be thought to be little interfered with by any
of these barriers, but, as Dr. Wallace has shown, they are almost as
much affected by them in their distribution as mammals are.
This then is the ordinary migratory distribution. Periodic distribution
obtains with migratory birds and fishes. The annual flight of swallows
to their northern summer residence comes under the heading of periodic
migration or distribution, but apart from this, the swallow must seek to
extend its range by the ordinary method, like every other animal.
Similarly, the herring migrates periodically into shallow water to
spawn, only to return again to its deeper home, where, as its numbers
increase, there must be a tendency to spread. We have in these cases,
therefore, both a periodic and an ordinary movement of migration.
Now, in studying the composition of a fauna, and especially its origin,
it is of the utmost importance to be able to determine approximately the
percentage of accidental arrivals and of the ordinary migrants--that is
to say, of those which have reached the country owing to accidental
distribution, and of animals which have adopted the usual course of
migration. It is of all the more import to review this subject of
accidental, or, as Darwin called it, "the occasional means" of
distribution, as both he and Dr. Wallace have, I venture to think,
somewhat over-estimated its significance. No one doubts that accidental
transportal takes place, but the question is whether the accidentally
transported animals arrive living and reach a spot where suitable food
is procurable, and whether they are able to propagate their own species
in the new locality. For it must be clear to anybody that the accidental
transportal of a beetle or of a snail to a new country cannot affect its
fauna or add one permanent member to it unless all these conditions are
fulfilled. As a matter of fact, only exceedingly few instances are on
record of man having witnessed, for example, the accidental transportal
across the sea to an island of a live animal.
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