The _variability of species_ has also been the subject of animated
discussion among naturalists. The memorable contention which arose
upon this subject between Cuvier and Geoffroy is not yet forgotten,
a struggle considered by Goethe as more important than the gravest
political events. In the present day a school to which many of the
most illustrious names in England, Germany and elsewhere belong,
has taken up, with certain modifications, the ideas of Lamarck
and Geoffroy; it gives support to them from retaining the term
_variability of species_.
There is a grave confusion of words in this formula. Lamarck,
Geoffroy, Darwin and his school, consider the species not only as
_variable_ but as _transmutable_. The specific types are not merely
_modified_, they are _replaced_ by new types. _Variation_ is in
their estimation only a phase of the very different phenomena of
_transmutation_.
I shall discuss these theories presently. I shall now confine
myself to the remark that _true variability_, admitted even by the
defenders of _dogmatic invariability_, by Blainville, for example,
a variability which I fully accept, has nothing in common with the
_transmutability_ of Lamarck, Geoffroy and Darwin. Let us briefly
determine the limits of this variability.
III. When an individual trait is exaggerated and passes a limit
always very loosely defined, it constitutes an exceptional character
which clearly distinguishes the individual affected by it from all
those most nearly resembling it. This individual constitutes a
_Variety_.
The same term must be applied to all those individuals, which, like
certain plants reproduced by slips, grafts, or shoots, derive their
origin from the first exceptional individual, without having the
power of transmitting their distinctive characters by means of normal
generation. I borrow from M. Chevreul a curious example of these
_multiple varieties_. In 1803 or 1805, M. Descemet discovered in his
garden at Saint Denis, in the midst of a bed of acacias (_Robinia
pseudo-acacia_) an individual without thorns which he describes
under the epithet _spectabilis_. It is to the multiplication of
this individual by the art of the gardener that all the _thornless
acacias_, now distributed over every part of the globe, owe their
origin. Now these individuals produce seeds, but if the seeds are
sown they only yield _thorny acacias_. The _acacia spectabilis_ has
remained a _Variety_.
The latter may then be defined as:—“An individual or a number of
individuals belonging to the same sexual generation, which is
distinguished from the other representatives of the same species by
one or several exceptional characters.”
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