The Kansas University Science Bulletin (Vol. I, No. 1)Various
Science
The Kansas University Science Bulletin (Vol. I, No. 1)
Various
Science -- Periodicals
Professor Todd’s theory in regard to the pollination of this plant is
as follows: “The weight of the bee so springs down the flower, that
it is quite difficult, on account of the large, flexible corolla,
to see just what is done, but repeated observations led me, quite
satisfactorily, to this conclusion. The bee seeks the pollen—for the
flowers have neither nectar nor odor—and this she uniformly gets from
the four shorter stamens; never, so far as I could determine, from the
larger one. This she does by seizing each one, near its base, between
her mandibles, and with a sort of milking motion crowds the pollen out
of the terminal pores; meanwhile, by the movements of her feet, the
larger stamen is repeatedly sprung backwards, and as often throws a
cloud of pollen on one side of her body; this in a right-handed flower.
When she passes to a left-handed flower, which, as was explained
above, is very likely not to be on the same plant, the pollen is
carried directly to the pistil of that flower, and so on. We have here,
therefore, a novel apparatus for cross-fertilization, quite distinct
from those that have been most commonly noticed.”
A considerable quantity of pollen may be thrown from the terminal pores
of the large stamen upon tapping it. It thus seems quite possible that
some pollen is thrown upon the side of the insect, as described by
Professor Todd. All the meaning of Mr. Meehan’s[K] statement is not
clear to the writers, but he says, in speaking of Professor Todd’s
results: “In regard to the manner in which the pollen is extracted, he
found that ‘this she does by seizing each anther near its base between
her mandibles, and, with a sort of milking motion, crowds the pollen
out of the terminal pores.’ If this were the general way, there would
be no necessity for any pollen being ejected from the long stamens,
for the stigma would surely receive some during the ‘milking’ process;
and the pore at the apex in the long anther is beyond the line of the
stigma, so that on ejection from the pore the pollen would go still
farther beyond.”
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