The Last Link: Our Present Knowledge of the Descent of ManHaeckel, Ernst
Science
The Last Link: Our Present Knowledge of the Descent of Man
Haeckel, Ernst
Human beings -- Origin
A. GAUDRY: 'Animaux fossiles et géologie de l'Attique.'
1862.--'Le Dryopithèque.' _Mém. Soc. géol. de France_:
'Paléontologie.' 1890.
O. MARSH: 'Introduction and Succession of Vertebrate Life in
America.' Address, Amer. Assoc. Adv. Sci., Nashville, 1887.
H. F. OSBORN: 'The Rise of the Mammalia in North America.'
Address, Amer. Assoc. Adv. Sci., Madison, 1893.
L. RUETIMEYER: 'Ueber die Herkunft unserer Thierwelt,' Basel,
1867.
C. S. FORSYTH MAJOR: 'Fossil Monkeys from Madagascar.'
_Geological Magazine_, 1896.
M. SCHLOSSER: 'Ueber die Beziehungen der ausgestorbenen
Saeugethierfaunen und ihr Verhaeltniss zur Saeugethierfauna der
Gegenwart.' Biolog. Centralblatt, 1888.
The most important difference between the two groups of existing
monkeys is indicated by their dentition. Adult man possesses, like
all the other Catarrhine Simiæ, thirty-two teeth, whilst the American
monkeys (the Platyrrhinæ) have thirty-six teeth--namely, one pair of
premolars more in the upper and lower jaws. Comparative odontology
leads us to the phylogenetic conclusion that this number has been
produced by reduction from a still older form with forty-four teeth.
This typical dental formula (three incisors, one canine, four
premolars, and three molars, in each half-jaw) is common to all those
most important older mammals which in the beginning of the Eocene
period constituted the four large groups of Lemuravida, Condylarthra,
Esthonychida, and Ictopsida. These are the four ancestral groups
of the four main orders of Placentalia--namely, of the Primates,
Ungulata, Rodentia, and Carnassia. They seem to be so closely related
by their primitive organization that they may be united in one common
super-order, Prochoriata.
With a considerable degree of probability, we are led to formulate
the further hypothesis that all the orders of Placentalia--from the
lowest Prochoriata upwards to man--have descended from some unknown
common ancestor living in the Cretaceous period, and that this oldest
placental form originated from some Jurassic group of marsupials.
Among these numerous fossil Lemures which have been discovered within
the last twenty years, there exist, indeed, all the connecting forms
of the older series of Primates, all the 'missing links' sought for by
comparative odontology.
The oldest Lemures of the tertiary age are the Eocene Pachylemures,
or Hyopsodina. They possess the complete dentition of the
Prochoriata--namely, forty-four teeth (3.1.4.3/3.1.4.3). Then follow
the Eocene Palæolemures, or Adapida, with forty teeth, they having lost
one pair of incisors in each jaw. To these are attached the younger
Autolemures, or Stenopida, with thirty-six teeth, they thus possessing
already the same dentition as the Platyrrhinæ. The characteristic
dentition of the Catarrhinæ is derived from this formula by the loss of
another premolar.
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