The Last Link: Our Present Knowledge of the Descent of ManHaeckel, Ernst
Science
The Last Link: Our Present Knowledge of the Descent of Man
Haeckel, Ernst
Human beings -- Origin
Man agrees in all these respects with the other vertebrates, and must
have descended with them from the same common root.
Particularly obscure is that part of our phylogeny which extends from
the Gastræa to Amphioxus. The morphological importance of this last
small creature had been perceived by Johannes Mueller, who in 1842
gave the first accurate description of it. It would not, of course, be
correct to proclaim the modern Amphioxus the common ancestor of all the
vertebrates; but he must be regarded as closely related to them, and
as the only survivor of the whole class of Acrania. If the Amphioxidæ
had through some unfortunate accident become extinct, we should not
have been able to gain anything like a positive glimpse at our most
remote vertebrate ancestor. On the one hand, Amphioxus is closely
connected with the early larva of the Cyclostomes, which are the
oldest Craniota, and the pre-Silurian ancestors of the fishes. On the
other hand, the ontogeny of Amphioxus is in harmony with that of the
Ascidians, and if this agreement is not merely coincidental, but due to
relationship, we are justified in reconstructing for both Ascidians
and Amphioxus one common ancestral group of chordate animals, the
hypothetical _Prochordonia_. The modern Copelata give us a remote idea
of their structure. The curious Balanoglossus, the only living form of
Enteropneusta, seems to connect these Prochordonia with the Nemertina
and other Vermalia, which we unite in one large class--Frontonia.
No doubt these pre-Cambrian Vermalia, and the common root of all
Metazoa, the Gastræades, were connected during the Laurentian period
by a long chain of intermediate forms, and probably among these
were some older forms of Rotatoria and Turbellaria; but at present
it is not possible to fill this wide gap with hypotheses that are
satisfactory, and we have to admit that here indeed are many missing
links in the older history of the Invertebrata. Still, every zoologist
who is convinced of the truth of transformism, and is accustomed to
phylogenetic speculations, knows very well that their results are most
unequal, often incomplete.
III.
Let us now recapitulate the ancestral chain of man, as it is set forth
in the accompanying diagram (p. 55), which represents our present
knowledge of our descent. For simplicity's sake the many side-issues
or branches which lead to groups not in the main line of our descent
have been left out, or have been indicated merely. Many of the stages
are of course hypothetical, arrived at by the study of comparative
anatomy and ontogeny; but an example for each of them has been taken
from those living or fossil creatures which seem to be their nearest
representatives.
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