The Last Link: Our Present Knowledge of the Descent of ManHaeckel, Ernst
Science
The Last Link: Our Present Knowledge of the Descent of Man
Haeckel, Ernst
Human beings -- Origin
4. The morula of most animals further changes into a _Blastula_, a
hollow ball filled with fluid, the wall being formed by a single layer
of cells, the blastoderm or germinal layer. This modification is
brought about by the action of the cells--they conveying nourishing
fluid into the interior of the whole cell colony and thereby
being themselves forced towards the surface. The Blastula of most
Invertebrata, and even that of Amphioxus, is possessed of fine ciliæ,
or hair-like processes, the vibrating motion of which causes the whole
organism to rotate and advance in the water. Living representatives of
such Blastæads, namely, globular gelatinous colonies of cells enclosing
a cavity, are Volvox and Magosphæra.
5. The Blastula of most animals assumes a new larval form called
_Gastrula_, in which the essential characteristics are that a portion
of the blastoderm by invagination converts the Blastula into a cup
with double walls, enclosing a new cavity, the primitive gut. This
invagination or bulging-in obliterates the original inner cavity of
the Blastula. The outer layer of the Gastrula is the ectoderm, the
inner the endoderm; both pass into each other at the blastoporus, or
opening of the gut cavity. The Gastrula is a stage in the embryonic
development of the various great groups of animals, and some such
primitive form as ancestral to all Metazoa is thus indicated. This
hypothetical _Gastræa_ is still very essentially represented by the
lower Coelenterates--_e.g._, Olynthus, Hydra.
6. The sixth stage--that of the _Platodes_, or flat-worms--is very
hypothetical. They are bilateral gastræads, with a flattened oblong
body, furnished with ciliæ, with a primitive nervous system, simple
sensory and reproductive organs, but still without appendages, body
cavity, vent, and blood-vessels. The nearest living representatives of
such creatures are the acoelous Turbellarians--_e.g._, Convoluta, a
free-swimming, ciliated creature.
7. The next higher stage is represented by such low animals as the
_Gastrotricha_--_e.g._, Chætonotus among the Rotatoria, which differ
from the rhabdocoelous Turbellarians chiefly by the formation of
a vent and the beginnings of a coelom, or cavity, between gut and
body wall. The addition of a primitive vascular system and a pair of
nephridia, or excretory organs, is first met with in the _Nemertines_.
8. These, together with the _Enteropneusta_ (Balanoglossus), are
comprised under the name of Frontonia, or Rhynchelminthes, and form the
highest group of the Vermalia.
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