In this marvel, so severe and dignified in shape and colouring, one
point above all captivates me and gives me at last the solution of a
far-reaching problem. Are the front-legs furnished with a tarsus, yes
or no? This is the great business that makes me forget the jewel for
the sake of a structural detail. Let us then return to a subject that
excited me in my early days; for the answer has come at last, late, it
is true, but certain and indisputable.
By a very strange exception, the full-grown Sacred Beetle and his
congeners are without front tarsi; they lack on their fore-legs that
five-jointed finger which is the rule among the highest division of
Coleoptera, the Pentamera. The other limbs, on the contrary, follow the
common law and possess a very well-shaped tarsus. Is the formation of
the toothed armlets original or accidental?
At first sight, an accident seems probable enough. The Scarab is a
strenuous miner and a great pedestrian. Always in contact with the
rough soil, whether in walking or digging; used, moreover, for constant
leverage when the insect is rolling its ball backwards, the fore-legs
are much more exposed than the others to the danger of spraining and
twisting their delicate finger, of putting it out of joint, of losing
it entirely, from the very first moment when the work begins.
Lest this explanation should appeal to any of my readers, I will hasten
to undeceive them. The absence of the front fingers is not the result
of an accident. The proof of what I say lies here, under my eyes,
without the possibility of a rejoinder. I examine the nymph’s legs with
the magnifying-glass: those in front have not the least vestige of a
tarsus; the toothed limb ends bluntly, without a trace of a terminal
appendage. In the others, on the contrary, the tarsus is as distinct as
possible, notwithstanding the shapeless, gnarled condition due to the
swaddling-bands and the humours of the chrysalis state. It suggests a
finger swollen with chilblains.
If the evidence of the nymph were not sufficient, there would be that
of the perfect insect which, casting its rejected mummy-clothes and
moving for the first time in its shell, wields fingerless armlets. The
fact, therefore, is established for certain: the Sacred Beetle is born
maimed; his mutilation dates from his birth.
Public-domain text, read in full here on John Shaqi.
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