The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
The third section of the Reptantia, the _Anomura_, comprises forms in
which the abdomen is variously modified, being either bent upon itself
or, if extended, more or less soft and feebly armoured. The last pair of
legs are commonly reduced in size, and not used in walking. The members
of the four tribes composing the section differ widely in their general
appearance.
The _Galatheidea_ (Plate VI.) are small, flattened, lobster-like animals
which have the abdomen bent under the body. In one family
(_Porcellanidæ_) the animals have quite the appearance of little Crabs
(see Fig. 41, p. 113), but they may be distinguished from the true Crabs
(Brachyura) by the fact that there are only three pairs of walking legs
behind the great chelæ, the last pair of legs being very small and
carried folded up at the sides of the body, or even within the gill
chambers.
[Illustration: _PLATE VI_
_Munida rugosa._ BRITISH. (REDUCED)]
The _Thalassinidea_ are small lobster-like animals which burrow in sand
and mud, and have generally a more or less soft abdomen (see Fig. 38, p.
103).
[Illustration: _PLATE VII_
THE COMMON HERMIT-CRAB, _Eupagurus bernhardus_, IN THE SHELL OF A
WHELK (REDUCED)
(_From Brit. Mus. Guide_)]
The tribe _Paguridea_ includes the Hermit Crabs (_Paguridæ_) and their
allies. The typical Hermit Crabs (Plate VII.), which are familiar
objects in seaside rock-pools, live in the empty shells of Whelks and
other Gasteropod Molluscs, which they carry about with them as portable
shelters. The structure of the animals is modified in adaptation to this
curious habit. The abdomen, which is protected during life by the
borrowed shell, is soft and unarmoured, and is spirally twisted. The
swimmerets, which have only the function of carrying the eggs in the
female, are much reduced, and are usually present only on one side of
the body. The uropods no longer form a tail-fan, but are adapted for
firmly wedging the hind part of the body into the coils of the shell.
One of the chelipeds is much larger than the other, and serves to block
up the opening when the animal withdraws into its shelter. In tropical
countries certain Hermit Crabs (_Coenobitidæ_) have become adapted to a
life on land, and one of these, the well-known Coconut Crab, or Robber
Crab (_Birgus latro_), which is the largest species of the tribe, has
given up the habit of protecting itself with a shell, and its abdomen
has again acquired a strong armour on the upper side. The marine
_Lithodidæ_--to which the British Stone Crab, _Lithodes maia_ (Plate
VIII.) belongs--seem at first sight to have little resemblance to the
Hermit Crabs, for they have the abdomen very small, and tucked up under
the body as in the true Crabs. Like the Porcellanidæ, mentioned above,
however, the Lithodidæ have only three pairs of walking legs behind the
chelipeds, the last pair being feeble and usually folded out of sight
within the gill chambers.
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